Thick-billed Flowerpecker
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The thick-billed flowerpecker (''Pachyglossa agilis'') is a tiny
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the
flowerpecker The flowerpeckers are a family (biology), family, Dicaeidae, of passerine birds. The family comprises three genera, ''Dicaeum'', ''Prionochilus'' and ''Pachyglossa'', with 56 species in total. The family has sometimes been included in an enlarge ...
group. They feed predominantly on fruits and are active birds that are mainly seen in the tops of trees in forests. It is a resident bird with a wide distribution across tropical southern
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
from
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
east to
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
and
Timor Timor (, , ) is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is Indonesia–Timor-Leste border, divided between the sovereign states of Timor-Leste in the eastern part and Indonesia in the ...
with several populations recognized as subspecies some of which are sometimes treated as full species. This species was formerly placed in the genus ''
Dicaeum ''Dicaeum'' is a genus of birds in the flowerpecker family Dicaeidae, a group of passerines tropical southern Asia and Australasia from India east to the Philippines and south to Australia. Within the family Dicaeidae the genus ''Dicaeum'' is ...
.


Taxonomy

The thick-billed flowerpecker was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
in 1833 by the English ornithologist
Samuel Tickell Colonel Samuel Richard Tickell (19 August 1811 – 20 April 1875) was an English soldier, artist, linguist and ornithologist in India and Burma. Biography Tickell was born at Cuttack in India to Captain Samuel Tickell (of the 8th regiment of th ...
based on a specimen collected in the Barabhum and
Dhalbhum Dhalbhum was an estate in early modern period and British India, spread across the East Singhbhum district in present-day Jharkhand,Ghosh, Binoy, ''Paschim Banger Sanskriti'', (in Bengali), part II, 1978 edition, p. 56, Prakash Bhaban western ...
forests of the state of
Jharkhand Jharkhand (; ) is a States and union territories of India, state in East India, eastern India. The state shares its border with the states of West Bengal to the east, Chhattisgarh to the west, Uttar Pradesh to the northwest, Bihar to the north ...
in eastern India. He placed the new species with the finches in the genus ''
Fringilla The genus ''Fringilla'' is a small group of eight species of finches from the Old World. It is the only genus in the subfamily Fringillinae. Taxonomy The genus ''Fringilla'' was described in 1758 by the Swedish naturalist Carl Linnaeus in the 1 ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Fringilla agilis''. The specific epithet is
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
meaning "nimble" or "active". The thick-billed flowerpecker was formerly placed in the genus ''
Dicaeum ''Dicaeum'' is a genus of birds in the flowerpecker family Dicaeidae, a group of passerines tropical southern Asia and Australasia from India east to the Philippines and south to Australia. Within the family Dicaeidae the genus ''Dicaeum'' is ...
'' but when molecular phylogenetic studies found that this genus was
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
, several species including the thick-billed flowerpecker were moved to the resurrected genus ''
Pachyglossa ''Pachyglossa'' is one of the three genera of flowerpeckers that make up the family Dicaeidae. These species are found in montane areas of South and Southeast Asia. Taxonomy The genus ''Pachyglossa'' was introduced in 1843 by the English zoologi ...
'' that had been introduced in 1843 by
Edward Blyth Edward Blyth (23 December 1810 – 27 December 1873) was an English zoologist who worked for most of his life in India as a curator of zoology at the Asiatic Society, Royal Asiatic Society of Bengal in Calcutta. He set about updating the museum ...
. Eleven
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised: * ''P. a. agilis'' ( Tickell, 1833) – northeast Pakistan and peninsular India * ''P. a. zeylonica'' ( Whistler, 1944) – Sri Lanka * ''P. a. pallescens'' (
Riley Riley may refer to: Businesses * Riley (brand), British sporting goods brand founded in 1878 * Riley Motor, British motorcar and bicycle manufacturera 1890–1969 * Riley Technologies, American auto racing constructor and team, founded by Bob ...
, 1935) – Bangladesh to Myanmar, north Thailand, and north Vietnam * ''P. a. modesta'' (
Hume Hume most commonly refers to: * David Hume (1711–1776), Scottish philosopher Hume may also refer to: People * Hume (surname) * Hume (given name) * James Hume Nisbet (1849–1923), Scottish-born novelist and artist In fiction * Hume, t ...
, 1875) – south peninsular Thailand, Malay Peninsula, and
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
* ''P. a. atjehensis'' ( Delacour, 1946) –
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
* ''P. a. finschi'' (Bartels, MEG, 1914) –
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...
(rare) * ''P. a. tincta'' (
Mayr Mayr is a German surname. Notable people with the surname include: * Andrea Mayr (born 1979), Austrian female long-distance runner * Ernst Mayr (1904–2005), German American evolutionary biologist * Franz Xaver Mayr (1875–1965), Austrian gastro ...
, 1944) –
Sumba Sumba (; ), natively also spelt as Humba, Hubba, Suba, or Zuba (in Sumba languages) is an Indonesian island (part of the Lesser Sunda Archipelago group) located in the Eastern Indonesia and administratively part of the East Nusa Tenggara pro ...
and
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Administratively, it forms the largest island in the East Nusa Tenggara Province. The area is 14,250 km2. Including Komodo and Rinca islands ...
to
Alor Island Alor () is the largest island in the Alor Archipelago and is one of the 92 officially listed outlying islands of Indonesia. It is located at the eastern end of the Lesser Sunda Islands chain that runs through southeastern Indonesia, which fr ...
(central
Lesser Sunda Islands The Lesser Sunda Islands (, , ), now known as Nusa Tenggara Islands (, or "Southeast Islands"), are an archipelago in the Indonesian archipelago. Most of the Lesser Sunda Islands are located within the Wallacea region, except for the Bali pro ...
) * ''P. a. obsoleta'' ( Müller, S, 1843) –
Timor Timor (, , ) is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is Indonesia–Timor-Leste border, divided between the sovereign states of Timor-Leste in the eastern part and Indonesia in the ...
and
Wetar Wetar is a tropical island which belongs to the Indonesian province of Maluku and is the largest island of the Maluku Barat Daya Islands (literally ''Southwest Islands'') Regency of the Maluku Islands. It lies east of the Lesser Sunda Islands, ...
(east
Lesser Sunda Islands The Lesser Sunda Islands (, , ), now known as Nusa Tenggara Islands (, or "Southeast Islands"), are an archipelago in the Indonesian archipelago. Most of the Lesser Sunda Islands are located within the Wallacea region, except for the Bali pro ...
) * ''P. a. striatissima''
Parkes Parkes may refer to: * Sir Henry Parkes (1815–1896), Australian politician, one of the earliest and most prominent advocates for Australian federation Named for Henry Parkes * Parkes, New South Wales, a regional town * Parkes Observatory, a radi ...
, 1962 –
Luzon Luzon ( , ) is the largest and most populous List of islands in the Philippines, island in the Philippines. Located in the northern portion of the List of islands of the Philippines, Philippine archipelago, it is the economic and political ce ...
group (north Philippines) * ''P. a. aeruginosa'' ( Bourns &
Worcester Worcester may refer to: Places United Kingdom * Worcester, England, a city and the county town of Worcestershire in England ** Worcester (UK Parliament constituency), an area represented by a Member of Parliament * Worcester Park, London, Engl ...
, 1894) –
Mindoro Mindoro is the seventh largest and eighth-most populous island in the Philippines. With a total land area of 10,571 km2 ( 4,082 sq.mi ), it has a population of 1,408,454, as of the 2020 census. It is located off the southwestern coast of ...
, West Visayas and
Mindanao Mindanao ( ) is the List of islands of the Philippines, second-largest island in the Philippines, after Luzon, and List of islands by population, seventh-most populous island in the world. Located in the southern region of the archipelago, the ...
(south Philippines) * ''P. a. affinis'' ( Zimmer, JT, 1919) –
Palawan Palawan (, ), officially the Province of Palawan (; ), is an archipelagic province of the Philippines that is located in the region of Mimaropa. It is the largest province in the country in terms of total area of . The capital and largest c ...
(southwest Philippines) The subspecies ''P. a. aeruginosa'', ''P. a. striatissima'' and ''P. a. affinis'' have sometimes been considered as a separate species, the striped flowerpecker.


Description

This species flowerpecker is about 10 cm long and has a dark stout beak and short tail. They are dark grey brown above and dull greyish with diffuse streaking on light buffy underparts. The rump is slightly more olive in the nominate race. The bill is dark, somewhat stout and heavy and the iris is reddish. The sexes are not distinguishable in the field and the juvenile has a paler base to the mandible and less streaks on the underside. There are whitish spots at the tip of the tail feathers. The nominate race is found on the plains of the mainland of the Indian Subcontinent. The Sri Lankan population, ''zeylonense'' (=''zeylonicum'', ''zeylonica''), is smaller and darker above. and Subspecies ''modestum'' (including ''pallescens'') is found in northeastern India and extends into Burma. Several island forms have been described but some of them are only tentatively kept within this species. These include ''atjehense'' of Sumatra, ''finschi'' of western Java, ''tinctum'' of Sumba, Flores and Alor, ''obsoletum'' from Timor, ''striatissimum'', ''aeruginosum'' and ''affine'' from the Philippines. Several of these such as ''aeruginosum'' are considered as full species as they are reproductively isolated and distinct in morphology.


Behaviour and ecology

Like other flowerpeckers they feed mainly on berries, nectar but sometimes take insects. Many of the subspecies are found in dense lowland forests with the exception of the nominate race which is found mainly in cultivated areas or open forest. Unlike the
pale-billed flowerpecker The pale-billed flowerpecker or Tickell's flowerpecker (''Dicaeum erythrorhynchos'') is a tiny bird that feeds on nectar and berries, found in India, Sri Lanka, Bangladesh and western Myanmar. The bird is common especially in urban gardens with b ...
, it does not swallow the berries of ''
Loranthus ''Loranthus'' is a genus of parasitic plants that grow on the branches of woody trees. It belongs to the family Loranthaceae, the showy mistletoe family. In most earlier systematic treatments it contains all mistletoe species with bisexual flower ...
'' (some species now in genus '' Dendrophthoe'') and instead wipes the seeds on a branch and feeds on the epicarp. This makes it disperse the parasitic mistletoe locally unlike the other species. In Sri Lanka, they forage at a greater height in the canopy than the pale-billed flowerpecker. Foraging birds produce a ''spick'' call frequently and the tail is jerked from side to side when perched. When displaying the male twitters and flutters over the female. The song is rambling and is mixed with notes that resemble that of the
ashy prinia The ashy prinia or ashy wren-warbler (''Prinia socialis'') is a small warbler in the family Cisticolidae. This prinia is a resident breeder in the Indian subcontinent, ranging across most of India, Nepal, Bangladesh, eastern Pakistan, Bhutan, Sri ...
. The feathers of the crown are erected in display and the white bases of the feathers become visible as a crown stripe. The nest has been described as appearing camouflaged like a dry leaf. It is a pendant purse like structure made of cobwebs or fine plant fibres and is located from 3 to 15 metres high suspended from a thin horizontal branch. Said to frequently nest among the nests of ''
Oecophylla smaragdina ''Oecophylla smaragdina'' (common names include Asian weaver ant, weaver ant, green ant, green tree ant, and orange gaster) is a species of tree, arboreal ant found in tropical Asia and Australia. These ants form colonies with multiple nests in ...
'' ants. The breeding season in southern India is December to March. Both male and female participate in nest building. The usual clutch is about 3 eggs but can vary from 2 to 4. The incubation period is around 13 days and the chick takes around 18 days to fledge. File:Thick-billed Flowerpecker (Dicaeum agile) on Helicteres isora W2 IMG 1379.jpg, ''Dicaeum agile agile'' on
Helicteres isora ''Helicteres isora'', sometimes called the Indian screw tree, is a small tree or large shrub found in southern Asia and northern Oceania. It is usually assigned to the family Malvaceae, but it is sometimes assigned to the family Sterculiaceae. Th ...
in Narsapur,
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
. File:DicaeumAgileWolf.jpg, Illustration of nest by
Joseph Wolf Joseph Wolf (22 January 1820 – 20 April 1899) was a German artist who specialized in natural history illustration. He moved to the British Museum in 1848 and became the preferred illustrator for explorers and naturalists including David Livin ...
Thick-billed flowerpecker - Timbi Lake, Vadodara, Gujarat 2023-12-03.jpg, ''Dicaeum agile agile'' from Gujarat


References


External links


Photographs and videos
{{Taxonbar, from=Q73848 thick-billed flowerpecker Birds of Indomalaya thick-billed flowerpecker