
Thallus (: thalli), from
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
ized
Greek
Greek may refer to:
Anything of, from, or related to Greece, a country in Southern Europe:
*Greeks, an ethnic group
*Greek language, a branch of the Indo-European language family
**Proto-Greek language, the assumed last common ancestor of all kno ...
(), meaning "a green
shoot
Shoot most commonly refers to:
* Shoot (botany), an immature plant or portion of a plant
* Shooting, the firing of projectile weapons
* Photo shoot, a photography session; an event wherein a photographer takes photographs
Shoot may also refer t ...
" or "
twig
A twig is a thin, often short, branch of a tree or Bush (plant), bush.
The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away. The color, texture, and patterning of the t ...
", is the vegetative tissue of some
organism
An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have be ...
s in diverse groups such as
algae
Algae ( , ; : alga ) is an informal term for any organisms of a large and diverse group of photosynthesis, photosynthetic organisms that are not plants, and includes species from multiple distinct clades. Such organisms range from unicellular ...
,
fungi
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
, some
liverworts
Liverworts are a group of non-vascular plant, non-vascular embryophyte, land plants forming the division Marchantiophyta (). They may also be referred to as hepatics. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in wh ...
,
lichen
A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
s, and the
Myxogastria
Myxogastria/Myxogastrea (myxogastrids, ICZN) or Myxomycetes ( ICN) is a class of slime molds that contains 5 orders, 14 families, 62 genera, and 888 species. They are colloquially known as the ''plasmodial'' or ''acellula ...
. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the
tissues into
organs
In a multicellular organism, an organ is a collection of tissues joined in a structural unit to serve a common function. In the hierarchy of life, an organ lies between tissue and an organ system. Tissues are formed from same type cells to a ...
. Many of these organisms were previously known as the
thallophyte
Thallophytes (Thallophyta or Thallobionta) are a polyphyletic group of non-motile organisms traditionally described as "thalloid plants", "relatively simple plants" or " lower plants". They form a division of kingdom Plantae that include lichens ...
s, a
polyphyletic
A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...
group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thalloidal, thalliform, thalline, or thallose.
Even though thalli do not have organized and distinct parts (
leave
Leave may refer to:
* Permission (disambiguation)
** Permitted absence from work
*** Leave of absence, a period of time that one is to be away from one's primary job while maintaining the status of employee
*** Annual leave, allowance of time awa ...
s,
root
In vascular plants, the roots are the plant organ, organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often bel ...
s, and
stems) as do the
vascular plant
Vascular plants (), also called tracheophytes (, ) or collectively tracheophyta (; ), are plants that have lignin, lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified Ti ...
s, they may have
analogous
Analogy is a comparison or correspondence between two things (or two groups of things) because of a third element that they are considered to share.
In logic, it is an inference or an argument from one particular to another particular, as oppose ...
structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has
vascular tissue
Vascular tissue is a complex transporting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. T ...
. In exceptional cases such as the
Lemnoideae
Lemnoideae is a subfamily of flowering aquatic plants, known as duckweeds, water lentils, or water lenses. They float on or just beneath the surface of still or slow-moving bodies of fresh water and wetlands. Also known as bayroot, they arose fr ...
, where the structure of a vascular plant is in fact thallus-like, it is referred to as having a thalloid structure, or sometimes as a thalloid.
Although a thallus is largely undifferentiated in terms of its anatomy, there can be visible differences and functional differences. A
kelp
Kelps are large brown algae or seaweeds that make up the order (biology), order Laminariales. There are about 30 different genus, genera. Despite its appearance and use of photosynthesis in chloroplasts, kelp is technically not a plant but a str ...
, for example, may have its thallus divided into three regions. The parts of a kelp thallus include the holdfast (anchor), stipe (supports the blades) and the blades (for photosynthesis).
The thallus of a
fungus
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
is usually called a
mycelium
Mycelium (: mycelia) is a root-like structure of a fungus consisting of a mass of branching, thread-like hyphae. Its normal form is that of branched, slender, entangled, anastomosing, hyaline threads. Fungal colonies composed of mycelium are fo ...
. The term thallus is also commonly used to refer to the
vegetative body of a lichen. In
seaweed
Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of ''Rhodophyta'' (red), '' Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ...
, thallus is sometimes also called '
frond
A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...
'.
The
gametophyte
A gametophyte () is one of the two alternating multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the se ...
of some non-thallophyte plants –
clubmoss
Lycopodiopsida is a class of vascular plants also known as lycopsids, lycopods, or lycophytes. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves ...
es,
horsetail
''Equisetum'' (; horsetail) is the only living genus in Equisetaceae, a family of vascular plants that reproduce by spores rather than seeds.
''Equisetum'' is a "living fossil", the only living genus of the entire subclass Equisetidae, which ...
s, and
fern
The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...
s is termed "
prothallus
A prothallus, or prothallium, (from Latin ''pro'' = forwards and Greek ''θαλλος'' (''thallos'') = twig) is usually the gametophyte stage in the life of a fern or other pteridophyte. Occasionally the term is also used to describe the young ...
".
See also
*
Homothallism In fungi and algae, homothallism refers to the condition in which a single individual or thallus carries the genetic determinants (i.e., both mating types or sexes) required to undergo sexual reproduction without the need for a distinct mating partn ...
*
Heterothallism
Heterothallic species have sexes that reside in different individuals. The term is applied particularly to distinguish heterothallic fungi, which require two compatible partners to produce sexual spores, from homothallic ones, which are capable ...
* ''
Wengania''
References
{{Reflist
Plant morphology
Fungal morphology and anatomy