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''Terminonaris'' is a genus of extinct
pholidosaurid Pholidosauridae is an extinct family of aquatic neosuchian mesoeucrocodylian crocodylomorphs. Fossils have been found in Europe (Denmark, England, France, Germany, Spain and Sweden), Africa (Algeria, Niger, Mali, Morocco and Tunisia), North Amer ...
crocodyliforms that lived during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
( Cenomanian and
Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded ...
Wu X-C, Russell AP, & Cumbaa SL. 2001. ''Terminonaris'' (Archosauria: Crocodyliformes): new material from Saskatchewan, Canada, and comments on its phylogenetic relationships. ''Journal of Vertebrate Paleontology'' 21(3):492-514.Shimada, K., and Parris, D.C., 2007. A long-snouted Late Cretaceous crocodyliform, ''Terminonaris'' cf. ''T. browni'', from the Carlile Shale (Turonian) of Kansas. ''Transactions of the Kansas Academy of Science'' 110(1):107-115.) epoch. The name means: ''“enlarged snout or nose”'' at the front of the skull. ''Terminonaris'' is an early crocodile, within a subgroup called Mesoeucrocodylia. Its remains have only been found in North America and
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
. Originally known under the generic name ''Teleorhinus'', it was once believed to be a teleosaurid (a family of marine gavial-like
thalattosuchia Thalattosuchia is a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of C ...
ns). Both prehistoric crocodiles such as ''Terminonaris'', as well as modern
crocodile Crocodiles (family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to include all extant ...
s, belong to the same group called
crocodyliformes Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only ps ...
, although modern crocodiles have specific features that indicate they are distant relatives of this species and members of the subgroup
Eusuchia Eusuchia is a clade of crocodylomorphs that first appeared in the Early Cretaceous with '' Hylaeochampsa''. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-T extinction. Since the other two clades ...
.


Description

''Terminonaris'' was a predator that could reach a length of about 6 m (19.7 ft). The skull of the largest individual is 98.3 cm (38.7 in) long. The elongated snout and the long, relatively thin teeth suggest that ''Terminonaris'' was a hunter of fish, small mammals and possibly small dinosaurs.


Habitat

''Terminonaris'' inhabited the
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient se ...
, an inland sea that stretched from the
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of the United S ...
to the
Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five major oceans. It spans an area of approximately and is known as the coldest of all the oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, ...
during Late Cretaceous time. The creature probably kept close to the shore, venturing into the seaway's warm salty water only to hunt for fish. Like modern crocodiles and alligators, ''Terminonaris'' would have eaten whatever it could catch, including small mammals and dinosaurs.


Specimens

Seven fossil specimens of ''Terminonaris'' have been discovered and all are from the Northern Hemisphere: six from North America and one from Europe. Scientists originally thought that ''Terminonaris'' originated in Europe, and then migrated across the Atlantic Ocean and dispersed throughout North America. However, it now seems more likely that ''Terminonaris'' originated in Texas and then dispersed northwar

All of the ''Terminonaris'' fossils from North America have been found along the shores of the former Western Interior Seaway. A Kansas specimen is the most recent, having lived about 91 million years ago, while those from Saskatchewan and Montana date from around 93 million years ago. The ''Terminonaris'' from Germany is believed to be from 94 million years ago. The oldest, from Texas, lived about 96 million years ago.


Big Bert

Big Bert is currently the world's most complete skeleton of ''Terminonaris robusta''. His bones were discovered in 1991 in an
outcrop An outcrop or rocky outcrop is a visible exposure of bedrock or ancient superficial deposits on the surface of the Earth. Features Outcrops do not cover the majority of the Earth's land surface because in most places the bedrock or superficia ...
of the
Favel Formation The Favel Formation is a stratigraphic unit of Late Cretaceous ( late Cenomanian to middle Turonian) age. It is present in southern Manitoba and southeastern Saskatchewan, and consists primarily of calcareous shale. It was named for the Favel Rive ...
along the banks of the Carrot River in the Pasquia Hills of
Saskatchewan Saskatchewan ( ; ) is a province in western Canada, bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and on the south by the U.S. states of Montana and North ...
.Cumbaa, S.L. and Tokaryk, T.S. 1999. Recent discoveries of Cretaceous marine vertebrates on the eastern margins of the Western Interior Seaway. In: Summary of Investigations 1999, Vol. 1, Saskatchewan Geological Survey, Sask. Energy Mines, Miscellaneous Report 99-4.1, p. 57-63. They were encased in an olive-black shale that smelled of oil. The discovery was made by
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of foss ...
Tim Tokaryk from the
Royal Saskatchewan Museum The Royal Saskatchewan Museum (RSM) is a Canadian natural history museum in Regina, Saskatchewan. Founded in 1906, it is the first museum in Saskatchewan, and the first provincial museum in the three Prairie Provinces. The institution was forme ...
, guided by local fossil hunter Dickson Hardie. The skeleton was recovered in 1992 by a team from the Royal Saskatchewan Museum and the
Canadian Museum of Nature The Canadian Museum of Nature (french: Musée canadien de la nature; CMN) is a national natural history museum based in Canada's National Capital Region. The museum's exhibitions and public programs are housed in the Victoria Memorial Museum ...
. They used explosives to dislodge some of the overburden that covered the skeleton. The area where Bert was discovered has yielded many fossils, including remains of birds, sharks, bony fish, dinosaurs, turtles, and
plesiosaurs The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
. At the time of discovery, Bert was the only one of his genus found in Canada. Big Bert is believed to have lived about 92 million years ago, at a time when the Carrot River area was near the eastern shoreline of the Western Interior Seaway. He was close to eight metres (25 feet) long and his skull was more than one metre (3 feet) long. His mouth was full of long skinny teeth, used for catching fish. To move in the water, Big Bert would tuck his small front feet close to his body and use his back feet and long tail to propel himself. Two models were cast from the fossilized bones of the original skeleton. One travels through Saskatchewan, visiting different museums. The second was installed at the Pasquia Regional Park in the Dickson Hardie Interpretive Centre, near Bert's Carrot River home.


Texas Terminonaris

In 2005 an amateur fossil enthusiast and rural mail carrier named Brian Condon discovered a 96 million year old ''Terminonaris'' while fossil-hunting close to his home on Lake Lewisville near
Dallas, Texas Dallas () is the third largest city in Texas and the largest city in the Dallas–Fort Worth metroplex, the fourth-largest metropolitan area in the United States at 7.5 million people. It is the largest city in and seat of Dallas County wi ...
. The fossils were located in an outcrop of the
Woodbine Formation The Woodbine Group is a geological formation in east Texas whose strata date back to the Early to Middle Cenomanian age of the Late Cretaceous. It is the producing formation of the giant East Texas Oil Field (also known as the "Black Giant") fro ...
. Mr. Condon donated the fossils to Southern Methodist University's Shuler Museum of Paleontology. They turned out to be the oldest examples of ''Terminonaris'', as well as the southernmost found to date.


References


External links


''Terminonaris''
in the
Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) originated in the NCEAS-funded Phanerozoic Marine Paleo ...

Pasquia Regional Park
{{Taxonbar, from=Q1027677 Late Cretaceous crocodylomorphs of North America Prehistoric pseudosuchian genera Prehistoric marine crocodylomorphs Late Cretaceous reptiles of North America Thalattosuchians