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The Terek sandpiper (''Xenus cinereus'') is a small migratory
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Si ...
wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
and is the only member of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''Xenus''. It is named after the
Terek River The Terek (; , Tiyrk; , Tərč; , ; , ; , ''Terk''; , ; , ) is a major river in the Northern Caucasus. It originates in the Mtskheta-Mtianeti region of Georgia and flows through North Caucasus region of Russia into the Caspian Sea The C ...
which flows into the west of the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the List of lakes by area, world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad s ...
, as it was first observed around this area.


Taxonomy

The Terek sandpiper was formally described and illustrated in 1775 by the German naturalist
Johann Anton Güldenstädt Johann Anton Güldenstädt (26 April 1745 in Riga, Latvia – 23 March 1781 in St. Petersburg, Russia) was a Baltic German naturalist and explorer in Russian service. Güldenstädt lost both his parents early, and from 1763 onwards studied ph ...
under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Scolopax cinerea''. He reported that he had seen pairs breeding at the mouth of the
Terek River The Terek (; , Tiyrk; , Tərč; , ; , ; , ''Terk''; , ; , ) is a major river in the Northern Caucasus. It originates in the Mtskheta-Mtianeti region of Georgia and flows through North Caucasus region of Russia into the Caspian Sea The C ...
where it flows into the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the List of lakes by area, world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad s ...
. It is now the only species placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''Xenus'' that was introduced in 1829 by the German naturalist
Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kau ...
. The genus name ''Xenus'' is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
''xenos'' meaning "stranger"; the specific epithet ''cinereus'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
for "ash-grey" from ''cinis, cineris'', "ashes". The species is considered to be
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
, no subspecies are recognised. Among the
Scolopacidae Sandpipers are a large family, Scolopacidae, of waders. They include many species called sandpipers, as well as those called by names such as curlew and snipe. The majority of these species eat small invertebrates picked out of the mud or soil. ...
, ''Xenus'' is part of the shank- tattler- phalarope clade and less closely related to the calidrid sandpipers. Based on the degree of
DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
divergence and putative shank and phalarope
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s from around the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
/
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
boundary some 23–22
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
, the Terek sandpiper presumably diverged from their relatives in the
Late Oligocene The Chattian is, in the geologic timescale, the younger of two ages or upper of two stages of the Oligocene Epoch/ Series. It spans the time between . The Chattian is preceded by the Rupelian and is followed by the Aquitanian (the lowest stag ...
. Given the numerous
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
fossils of the group found in
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
it is likely that the Terek sandpiper lineage originated there, possibly by being isolated as the remains of the Turgai Sea dried up, which happened just around this time.


Description

Slightly larger than the
common sandpiper The common sandpiper (''Actitis hypoleucos'') is a small Palearctic wader. This bird and its American sister species, the spotted sandpiper (''A. macularia''), make up the genus ''Actitis''. They are parapatric and replace each other geographical ...
at length, its long upcurved bill – somewhat reminiscent of an
avocet The four species of avocets are a genus, ''Recurvirostra'', of waders in the same avian family as the stilts. The genus name comes from Latin , 'curved backwards' and , 'bill'. The common name is thought to derive from the Italian ( Ferrarese) ...
's, but not as strongly curved – makes it very distinctive. As the scientific specific name implies, this wader has a grey back, face and breast in all
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
s; a white
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
may appear more or less distinct. The belly is whitish and the feet yellow; the bill has a yellowish base, with the rest being black. The call is a high whistle.


Distribution and ecology

This bird breeds near water in the
taiga Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces ...
from
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bo ...
through northern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
to the
Kolyma River The Kolyma ( rus, Колыма, p=kəlɨˈma; sah, Халыма, translit=Khalyma) is a river in northeastern Siberia, whose basin covers parts of the Sakha Republic, Chukotka Autonomous Okrug, and Magadan Oblast of Russia. The Kolyma is froze ...
, and
migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
south in winter to tropical coasts in east Africa, south Asia and Australia, usually preferring muddy areas. It is a rare vagrant in western Europe, and particularly in autumn it is sometimes seen passing through the Marianas on migration; on
Palau Palau,, officially the Republic of Palau and historically ''Belau'', ''Palaos'' or ''Pelew'', is an island country and microstate in the western Pacific. The nation has approximately 340 islands and connects the western chain of the Ca ...
, further off its usual migration route, it is decidedly uncommon on the other hand. Almost annually and apparently more and more often in recent times, a few birds stray to
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S ...
and the Aleutian and
Pribilof Islands The Pribilof Islands (formerly the Northern Fur Seal Islands; ale, Amiq, russian: Острова Прибылова, Ostrova Pribylova) are a group of four volcanic islands off the coast of mainland Alaska, in the Bering Sea, about north of ...
. Every few years, individual vagrants are recorded in the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bi ...
, where they arrive either as migrating birds from Africa, or as North American strays accompanying local waders south for winter. Such vagrants have been recorded as far south as
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
. The overall genetic variation in Terek sandpipers across their range is low, with some evidence of contractions followed by expansion. Although the geographically isolated Dniephe River population in Eastern Europe does show significant genetic differentiation. It feeds in a distinctive and very active way, chasing insects and other mobile prey, and sometimes then running to the water's edge to wash its catch. It lays three or four eggs in a lined ground scrape. The Terek sandpiper likes to associate with
ruddy turnstone The ruddy turnstone (''Arenaria interpres'') is a small cosmopolitan wading bird, one of two species of turnstone in the genus ''Arenaria''. It is now classified in the sandpiper family Scolopacidae but was formerly sometimes placed in the pl ...
s (''Arenaria interpres''), smallish calidrids, and ''
Charadrius '' Charadrius '' is a genus of plovers, a group of wading birds. The genus name ''Charadrius'' is a Late Latin word for a yellowish bird mentioned in the fourth-century Vulgate. They are found throughout the world. Many ''Charadrius'' species a ...
'' (but maybe not '' Pluvialis'')
plover Plovers ( , ) are a widely distributed group of wading birds belonging to the subfamily Charadriinae. Description There are about 66 species in the subfamily, most of them called "plover" or " dotterel". The closely related lapwing su ...
s; a vagrant bird at
Paraty Paraty (or Parati, ) is a preserved Portuguese colonial (1500–1822) and Brazilian Imperial (1822–1889) municipality with a population of about 43,000. Indeed, the name "Paraty" originates from the local Guaianá Indians' Indigenous Tupi l ...
(
Rio de Janeiro state Rio de Janeiro () is one of the 27 federative units of Brazil. It has the second largest economy of Brazil, with the largest being that of the state of São Paulo. The state, which has 8.2% of the Brazilian population, is responsible for 9.2% of ...
) was noted to pair up with a spotted sandpiper (''Actitis macularius''). This is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' (
AEWA The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA) is an independent international treaty developed under the auspices of the United Nations Environment Programme's Convent ...
) applies. Widespread and often quite commonly seen, the Terek sandpiper is not considered a threatened species by the IUCN.


Gallery

File:Terek Sandpiper, Little Stint, Marsh Sandpiper, C Redshank W2 IMG 6891.jpg, Terek sandpiper ''Xenus cinereus'',
little stint The little stint (''Calidris minuta'' or ''Erolia minuta''), is a very small wader. It breeds in arctic Europe and Asia, and is a long-distance migrant, wintering south to Africa and south Asia. It occasionally is a vagrant to North America ...
''Calidris minuta'' and
marsh sandpiper The marsh sandpiper (''Tringa stagnatilis'') is a small wader. It is a rather small shank, and breeds in open grassy steppe and taiga wetlands from easternmost Europe to the Russian Far East. The genus name ''Tringa'' is the New Latin name given ...
''Tringa stagnatilis'' in Krishna Wildlife Sanctuary,
Andhra Pradesh Andhra Pradesh (, abbr. AP) is a state in the south-eastern coastal region of India. It is the seventh-largest state by area covering an area of and tenth-most populous state with 49,386,799 inhabitants. It is bordered by Telangana to t ...
,
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
. File:Xenus cinereus eating ragworm.JPG, Eating ragworm in
Shōnai River The is a Class 1 river flowing through Gifu and Aichi prefectures in Japan. In Gifu Prefecture, it is also referred to as the Toki River (土岐川 ''Toki-gawa''); around the city of Kasugai in Aichi Prefecture, it is referred to as the Tam ...
, Japan. File:Xenus cinereus MWNH 0248.JPG, Egg, Collection Museum Wiesbaden File:Terek Sandpiper (Xenus cinereus) (8079446567).jpg, In flight File:Terek Sandpiper AMSM5480 TSAN.jpg, In Narara,
Jamnagar Jamnagar () is a city located on the western coast of India in the state of Gujarat of Saurashtra region. It is the administrative headquarters of the Jamnagar district and the fifth largest city in Gujarat. The city lies just to the south o ...
,
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
File:Terek Sandpiper AMSM5481.jpg, In Narara,
Jamnagar Jamnagar () is a city located on the western coast of India in the state of Gujarat of Saurashtra region. It is the administrative headquarters of the Jamnagar district and the fifth largest city in Gujarat. The city lies just to the south o ...
,
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...


References


External links


Terek sandpiper species text in The Atlas of Southern African Birds
* * * * * * * {{Taxonbar, from=Q18856 Terek sandpiper Birds of Russia Birds of Central Asia Terek sandpiper Terek sandpiper