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Temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
deciduous or temperate broadleaf forests are a variety of
temperate forest A temperate forest is a forest found between the tropical and boreal regions, located in the temperate zone. It is the second largest terrestrial biome, covering 25% of the world's forest area, only behind the boreal forest, which covers about 3 ...
'dominated' by
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
trees that lose their leaves each winter. They represent one of Earth's major
biome A biome () is a distinct geographical region with specific climate, vegetation, and animal life. It consists of a biological community that has formed in response to its physical environment and regional climate. In 1935, Tansley added the ...
s, making up 9.69% of global land area. These forests are found in areas with distinct seasonal variation that cycle through warm, moist summers, cold winters, and moderate fall and spring seasons. They are most commonly found in the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
, with particularly large regions in eastern
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
,
East Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
, and a large portion of
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
, though smaller regions of temperate deciduous forests are also located in South America. Examples of trees typically growing in the Northern Hemisphere's deciduous forests include
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
,
maple ''Acer'' is a genus of trees and shrubs commonly known as maples. The genus is placed in the soapberry family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated si ...
, basswood,
beech Beech (genus ''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to subtropical (accessory forest element) and temperate (as dominant element of Mesophyte, mesophytic forests) Eurasia and North America. There are 14 accepted ...
and
elm Elms are deciduous and semi-deciduous trees comprising the genus ''Ulmus'' in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical- montane regions of North America and Eurasia, ...
, while in the Southern Hemisphere, trees of the genus ''
Nothofagus ''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere, found across southern South America (Chile, Argentina) and east and southeast Australia, New Zealand, New Guin ...
'' dominate this type of forest. Temperate deciduous forests provide several unique
ecosystem service Ecosystem services are the various benefits that humans derive from ecosystems. The interconnected living and non-living components of the natural environment offer benefits such as pollination of crops, clean air and water, decomposition of wast ...
s, including
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s for diverse wildlife, and they face a set of natural and human-induced disturbances that regularly alter their structure.


Geography

Located below the northern
boreal forests Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. In North Ame ...
, temperate deciduous forests make up a significant portion of the land between the
Tropic of Cancer The Tropic of Cancer, also known as the Northern Tropic, is the Earth's northernmost circle of latitude where the Sun can be seen directly overhead. This occurs on the June solstice, when the Northern Hemisphere is tilted toward the Sun ...
(23\tfrac°N) and latitudes of 50° North, in addition to areas south of the
Tropic of Capricorn The Tropic of Capricorn (or the Southern Tropic) is the circle of latitude that contains the subsolar point at the December (or southern) solstice. It is thus the southernmost latitude where the Sun can be seen directly overhead. It also reach ...
(23\tfrac°S). Canada, the United States, China, and several European countries have the largest land area covered by temperate deciduous forests, with smaller portions present throughout South America, specifically Chile and Argentina.


Climate

Temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
conditions refer to the cycle through four distinct
season A season is a division of the year based on changes in weather, ecology, and the number of daylight hours in a given region. On Earth, seasons are the result of the axial parallelism of Earth's axial tilt, tilted orbit around the Sun. In temperat ...
s that occurs in areas between the polar regions and tropics. In these regions where temperate deciduous forest are found, warm and cold air circulation accounts for the biome's characteristic seasonal variation.


Temperature

The average annual temperature tends to be around 10 °Celsius, though this is dependent on the region. Due to shading from the canopy, the
microclimate A microclimate (or micro-climate) is a local set of atmosphere of Earth, atmospheric conditions that differ from those in the surrounding areas, often slightly but sometimes substantially. The term may refer to areas as small as a few square m ...
of temperate deciduous forests tends to be about 2.1 °Celsius cooler than the surroundings, whereas winter temperatures are from 0.4 to 0.9 °Celsius warmer within forests as a result of insulation from vegetation strata.


Precipitation

Annually, temperate deciduous forests experience approximately 750 to 1,500 millimeters of
precipitation In meteorology, precipitation is any product of the condensation of atmospheric water vapor that falls from clouds due to gravitational pull. The main forms of precipitation include drizzle, rain, rain and snow mixed ("sleet" in Commonwe ...
. As there is no distinct rainy season, precipitation is spread relatively evenly throughout the year. Snow makes up a portion of the precipitation present in temperate deciduous forests in the
winter Winter is the coldest and darkest season of the year in temperate and polar climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Dif ...
. Tree branches can intercept up to 80% of snowfall, affecting the amount of snow that ultimately reaches and melts on the forest floor.


Seasonal variation

A factor of temperate deciduous forests is their leaf loss during the transition from fall to winter, an adaptation that arose as a solution for the low sunlight conditions and bitter cold temperatures. In these forests, winter is a time of dormancy for plants, when broadleaf deciduous trees conserve energy and prevent water loss, and many animal species hibernate or migrate. Preceding winter is fruit-bearing autumn, a time when leaves change color to various shades of red, yellow, and orange as
chlorophyll Chlorophyll is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words (, "pale green") and (, "leaf"). Chlorophyll allows plants to absorb energy ...
breakdown gives rise to anthocyanin, carotene, and xanthophyl pigments. Besides the characteristic colorful autumns and leafless winters, temperate deciduous forests have a lengthy growing season during the spring and
summer Summer or summertime is the hottest and brightest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, daylight hours are the longest and darkness hours are the shortest, with day ...
months that tends to last anywhere from 120 to 250 days. Spring in temperate deciduous forests is a period of ground vegetation and seasonal herb growth, a process that starts early in the season before trees have regrown their leaves and when ample sunlight is available. Once a suitable temperature is reached in mid- to late spring, budding and flowering of tall deciduous trees also begins. In the summer, when fully-developed leaves occupy all trees, a moderately-dense canopy creates shade, increasing the humidity of forested areas.


Characteristics


Soil

Though there is latitudinal variation in
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
quality of temperate deciduous forests, with those at central latitudes having a higher soil productivity than those more north or south, soil in this biome is overall highly fertile. The fallen leaves from deciduous trees introduce
detritus In biology, detritus ( or ) is organic matter made up of the decomposition, decomposing remains of organisms and plants, and also of feces. Detritus usually hosts communities of microorganisms that colonize and decomposition, decompose (Reminera ...
to the
forest floor The forest floor, also called detritus or wikt:duff#Noun 2, duff, is the part of a forest ecosystem that mediates between the living, aboveground portion of the forest and the mineral soil, principally composed of dead and decaying plant matter ...
, increasing levels of nutrients and
organic matter Organic matter, organic material or natural organic matter is the large source of carbon-based compounds found within natural and engineered, terrestrial, and aquatic environments. It is matter composed of organic compounds that have come fro ...
in the soil. The high soil productivity of temperate deciduous forests puts them at a high risk of conversion to
agricultural land Agricultural land is typically land ''devoted to'' agriculture, the systematic and controlled use of other organism, forms of lifeparticularly the rearing of livestock and production of cropsto produce food for humans. It is generally synonymous ...
for human use.


Flora

Temperate deciduous forests are characterized by a variety of temperate deciduous tree species that vary based on region. Most tree species present in temperate deciduous forests are broadleaf trees that lose their leaves in the fall, though some
conifer Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
ous trees such as pines (''Pinus'') are present in northern temperate deciduous forests. Europe's temperate deciduous forests are rich with oaks of the genus ''Quercus'', European beech trees (''Fagus sylvatica''), and hornbeams ''(Fagus grandifolia''), while those in Asia tend to have maples of the genus ''Acer'', a variety of ash trees (''Fraxinus''), and basswoods (''Tilia''). Similarly to Asia, North American forests have maples, especially ''Acer saccharum,'' and basswoods, in addition to hickories ''(Carya'') and American chestnuts (''Castanea dentata''). Southern beech ''(Nothofagus)'' trees are prevalent in the temperate deciduous forests of South America. Elm trees (''Ulmus'') and willows (''Salix'') can also be found dispersed throughout the temperate deciduous forests of the world. While a wide variety of tree species can be found throughout the temperate deciduous forest biome, tree
species richness Species richness is the number of different species represented in an community (ecology), ecological community, landscape or region. Species richness is simply a count of species, and it does not take into account the Abundance (ecology), abunda ...
is typically moderate in each individual ecosystem, with only 3 to 4 tree species per square kilometer. Besides the old-growth trees that, with their domed tree crowns, form a
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
that lets little light filter through, a sub-canopy of
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s such as mountain laurel and azaleas is present. These other plant species found in the canopy layers below the 35- to 40-meter mature trees are either adapted to low-light conditions or follow a seasonal schedule of growth that allows them to thrive before the formation of the canopy from mid-spring through mid-fall.
Moss Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...
es and
lichen A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
s make up significant ground cover, though they are also found growing on trees.


Fauna

In addition to characteristic flora, temperate deciduous forests are home to several animal species that rely on the trees and other plant life for shelter and resources, such as
squirrel Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
s,
rabbit Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
s,
skunk Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or gi ...
s,
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s, mountain lions,
bobcat The bobcat (''Lynx rufus''), also known as the wildcat, bay lynx, or red lynx, is one of the four extant species within the medium-sized wild cat genus '' Lynx''. Native to North America, it ranges from southern Canada through most of the c ...
s, timber wolves, foxes, and black bears.
Deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
are also present in large populations, though they are clearing rather than true forest animals. Large deer populations have deleterious effects on tree regeneration overall, and grazing also has significant negative effects on the number and kind of herbaceous flowering plants. The continuous increase of deer populations and killing of top carnivores suggests that
overgrazing Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature ...
by deer will continue.


Ecosystem services

Temperate deciduous forests provide several provisioning, regulating, supporting, and cultural ecosystem services. With a higher
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
than boreal forests, temperate deciduous forests maintain their genetic diversity by providing the supporting service of habitat availability for a variety of plants and animal species dependent on shade. These forests play a role in the regulation of air and soil quality by preventing soil
erosion Erosion is the action of surface processes (such as Surface runoff, water flow or wind) that removes soil, Rock (geology), rock, or dissolved material from one location on the Earth's crust#Crust, Earth's crust and then sediment transport, tran ...
and
flood A flood is an overflow of water (list of non-water floods, or rarely other fluids) that submerges land that is usually dry. In the sense of "flowing water", the word may also be applied to the inflow of the tide. Floods are of significant con ...
ing, while also storing carbon in their soil. Provisioning services provided by temperate deciduous forests include access to sources of drinking water, oxygen, food,
timber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...
, and
biomass Biomass is a term used in several contexts: in the context of ecology it means living organisms, and in the context of bioenergy it means matter from recently living (but now dead) organisms. In the latter context, there are variations in how ...
. Humans depend on temperate deciduous forests for cultural services, using them as spaces for recreation and spiritual practices.


Disturbances

Natural disturbances cause regular renewal of temperate deciduous forests and create a healthy, heterogeneous environment with constantly changing structures and populations. Weather events like snow, storms, and wind can cause varying degrees of change to the structure of forest canopies, creating log habitats for small animals and spaces for less shade-tolerant species to grow where fallen trees once stood. Other
abiotic In biology and ecology, abiotic components or abiotic factors are non-living chemical and physical parts of the environment that affect living organisms and the functioning of ecosystems. Abiotic factors and the phenomena associated with them und ...
sources of disturbances to temperate deciduous forests include
drought A drought is a period of drier-than-normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, ...
s, waterlogging, and fires. Natural surface fire patterns are especially important in
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
reproduction. Biotic factors affecting forests take the form of fungal outbreaks in addition to mountain pine beetle and
bark beetle A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family (Curculionidae). Although th ...
infestations. These beetles are particularly prevalent in North America and kill trees by clogging their
vascular tissue Vascular tissue is a complex transporting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. T ...
. Temperate deciduous forests tend to be resilient after minor weather-related disturbances, though major insect infestations, widespread anthropogenic disturbances, and catastrophic weather events can cause century-long
succession Succession is the act or process of following in order or sequence. Governance and politics *Order of succession, in politics, the ascension to power by one ruler, official, or monarch after the death, resignation, or removal from office of ...
or even the permanent conversion of the forest into a
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
.


Climate change

Rising temperatures and increased dryness in temperate deciduous forests have been noted in recent years as the
climate change Present-day climate change includes both global warming—the ongoing increase in Global surface temperature, global average temperature—and its wider effects on Earth's climate system. Climate variability and change, Climate change in ...
s. As a result, temperate deciduous forests have been experiencing an earlier onset to spring, as well as a global increase in the frequency and intensity of disturbances. They have been experiencing lower ecological resilience in the face of increasing mega-fires, longer droughts, and severe storms. Damaged wood from increased storm disturbance events provides nesting habitats for beetles, concurrently increasing bark beetle damage. Forest cover decreases with continuous severe disturbances, causing
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
and lower biodiversity.


Human use and impact

Humans rely on
wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
from temperate deciduous forests for use in the
timber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...
industry as well as paper and
charcoal Charcoal is a lightweight black carbon residue produced by strongly heating wood (or other animal and plant materials) in minimal oxygen to remove all water and volatile constituents. In the traditional version of this pyrolysis process, ca ...
production.
Logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidder, skidding, on-site processing, and loading of trees or trunk (botany), logs onto logging truck, trucks During the European colonization of North America,
potash Potash ( ) includes various mined and manufactured salts that contain potassium in water- soluble form.
made from tree ashes was exported back to Europe as fertilizer. At this time in history,
clearcutting Clearcutting, clearfelling or clearcut logging is a forestry/logging practice in which most or all trees in an area are uniformly cut down. Along with Shelterwood cutting, shelterwood and Seed tree, seed tree harvests, it is used by foresters t ...
of the original temperate deciduous forests was also performed to make space for agricultural land use, so many forests now present are second-growth. Over 50% of temperate deciduous forests are affected by fragmentation, resulting in small fragments dissected by fields and roads; these islands of green often differ substantially from the original forests and cause challenges for species migration. Seminatural temperate deciduous forests with developed trail systems serve as sites for tourism and recreational activities, such as hiking and hunting. In addition to fragmentation, human use of land adjacent to temperate deciduous forests is associated with
pollution Pollution is the introduction of contaminants into the natural environment that cause harm. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the component ...
that can stunt the growth rate of trees.
Invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
that outcompete
native species In biogeography, a native species is indigenous to a given region or ecosystem if its presence in that region is the result of only local natural evolution (though often popularised as "with no human intervention") during history. The term is equi ...
and alter forest
nutrient cycle A nutrient cycle (or ecological recycling) is the movement and exchange of inorganic and organic matter back into the production of matter. Energy flow is a unidirectional and noncyclic pathway, whereas the movement of mineral nutrients is cyc ...
s, such as common buckthorn ''(Rhamnus cathartica)'', are also introduced by humans. The introduction of exotic diseases, especially, continues to be a threat to forest trees and, hence, the forest. Humans have also introduced
earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s in deciduous forests in Norh America, which has had a deep impact on the ecosystem and reduced biodiversity.Invasion of a deciduous forest by earthworms: Changes in soil chemistry, microflora, microarthropods and vegetation
/ref>


Conservation

A method for preserving temperate deciduous forests that has been used in the past is
fire suppression Fire suppression may refer to: * Firefighting * Fire suppression systems * Wildfire suppression Wildfire suppression is a range of firefighting tactics used to suppress wildfires. Firefighting efforts depend on many factors such as the availabl ...
. The process of preventing fires is associated with the build-up of biomass that, ultimately, increases the intensity of incidental fires. As an alternative, prescribed burning has been put into practice, in which regular, managed fires are administered to forest ecosystems to imitate the natural disturbances that play a significant role in preserving biodiversity. To combat the effects of
deforestation Deforestation or forest clearance is the removal and destruction of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. Ab ...
,
reforestation Reforestation is the practice of restoring previously existing forests and woodlands that have been destroyed or damaged. The prior forest destruction might have happened through deforestation, clearcutting or wildfires. Three important purpose ...
has been employed.


See also

*
Temperate coniferous forest Temperate coniferous forest is a terrestrial biome defined by the World Wide Fund for Nature. Temperate coniferous forests are found predominantly in areas with warm summers and cool winters, and vary in their kinds of plant life. In some, needle ...
*
Temperate broadleaf and mixed forest Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions. Thes ...
* International Year of Forests *
Old-growth forest An old-growth forest or primary forest is a forest that has developed over a long period of time without disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organization of the United Natio ...
* Tropical evergreen forest * Tropical deciduous forest * Wood-pasture hypothesis


References


External links

*
A map of biome distribution
(Temperate Deciduous Forest is in dark green) {{Biomes Deciduous Terrestrial biomes Forests Habitat