The term tarpan (''Equus ferus ferus'') refers to free-ranging horses of the Russian steppe from the 18th to the 20th century. It is generally unknown whether those horses represented genuine wild horses, feral domestic horses or hybrids. The last individual believed to be a tarpan died in captivity in the
Russian Empire
The Russian Empire was an empire and the final period of the List of Russian monarchs, Russian monarchy from 1721 to 1917, ruling across large parts of Eurasia. It succeeded the Tsardom of Russia following the Treaty of Nystad, which ended th ...
during 1909.
Beginning in the 1930s, several attempts were made to develop horses that looked like tarpans through
selective breeding
Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant ma ...
, called " breeding back" by advocates. The breeds that resulted included the Heck horse, the , and a derivation of the
Konik
The Konik or Polish Konik, pl, konik polski, is a Polish breed of pony. There are semi-feral populations in some regions. They are usually mouse dun or striped dun in color.
The Bilgoray, pl, konik biłgorajski, of south-eastern Poland is a ...
breed, all of which have a primitive appearance, particularly in having the
grullo
Grulla or grullo, also called blue dun, gray dun or mouse dun, is a equine coat color, color of horses in the dun (color), dun family, characterized by tan-gray or mouse-colored hairs on the body, often with shoulder and dorsal stripes and black ...
coat colour. Some of these horses are now commercially promoted as "tarpans", although such animals are only domestic breeds and not the wild animal themselves.
Name and etymology
The name "tarpan" or "tarpani" derives from a
Turkic language
The Turkic languages are a language family of over 35 documented languages, spoken by the Turkic peoples of Eurasia from Eastern Europe and Southern Europe to Central Asia, East Asia, North Asia (Siberia), and Western Asia. The Turkic languag ...
(
Kazakh
Kazakh, Qazaq or Kazakhstani may refer to:
* Someone or something related to Kazakhstan
*Kazakhs, an ethnic group
*Kazakh language
*The Kazakh Khanate
* Kazakh cuisine
* Qazakh Rayon, Azerbaijan
*Qazax, Azerbaijan
*Kazakh Uyezd, administrative dis ...
wild horse
The wild horse (''Equus ferus'') is a species of the genus ''Equus'', which includes as subspecies the modern domesticated horse (''Equus ferus caballus'') as well as the endangered Przewalski's horse (''Equus ferus przewalskii''). The Euro ...
". The
Tatars
The Tatars ()Tatar in the Collins English Dictionary is an umbrella term for different and the Cossacks distinguished the wild horse from the
feral horse
A feral horse is a free-roaming horse of domesticated stock. As such, a feral horse is not a wild animal in the sense of an animal without domesticated ancestors. However, some populations of feral horses are managed as wildlife, and these ...
; the latter was called ''Takja'' or ''Muzin''.
Taxonomy
The tarpan was first described by Samuel Gottlieb Gmelin in 1771; he had seen the animals in 1769 in the district of Bobrov, near
Voronezh
Voronezh ( rus, links=no, Воро́неж, p=vɐˈronʲɪʂ}) is a city and the administrative centre of Voronezh Oblast in southwestern Russia straddling the Voronezh River, located from where it flows into the Don River. The city sits on ...
. In 1784,
Pieter Boddaert
Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and naturalist.
Early life, family and education
Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D. at the Univers ...
named the species ''Equus ferus'', referring to Gmelin's description. Unaware of Boddaert's name, Otto Antonius published the name ''Equus gmelini'' in 1912, again referring to Gmelin's description. Since Antonius' name refers to the same description as Boddaert's it is a junior objective synonym. It is now thought that the domesticated
horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million ...
, named ''Equus caballus'' by
Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in 1758, is descended from the tarpan; indeed, many taxonomists consider them to belong to the same species. By a strict application of the rules of the
International Code of Zoological Nomenclature
The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the Int ...
, the tarpan ought to be named ''E. caballus'', or if considered a subspecies, ''E. caballus ferus''. However, biologists have generally ignored the letter of the rule and used ''E. ferus'' for the tarpan to avoid confusion with the domesticated subspecies.
It is debated if the small, free-roaming horses seen in the Russian steppes during 18th and 19th centuries and called "tarpan" were indeed wild, never-domesticated horses, hybrids of the
Przewalski's horse
Przewalski's horse (, , (Пржевальский ), ) (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered horse originally native to the steppes of Ce ...
and local domestic animals, or simply
feral horse
A feral horse is a free-roaming horse of domesticated stock. As such, a feral horse is not a wild animal in the sense of an animal without domesticated ancestors. However, some populations of feral horses are managed as wildlife, and these ...
s. Most studies have been based on only two preserved specimens and research to date has not positively linked the tarpan to Pleistocene or Holocene-era animals.
In 2003, the
International Commission on Zoological Nomenclature
The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries.
Orga ...
"conserved the usage of 17 specific names based on wild species, which are predated by or contemporary with those based on domestic forms", confirming ''E. ferus'' for the undomesticated wild horse. Taxonomists who consider the domestic horse a subspecies of the wild horse should use ''Equus ferus caballus''; the name ''Equus caballus'' remains available for the domestic horse where it is considered to be a separate species.
Appearance
Traditionally, two tarpan subtypes have been proposed, the forest tarpan and steppe tarpan, although there seem to be only minor differences in type. The general view is that there was only one subspecies, the tarpan, ''Equus ferus ferus''. The last individual, which died in captivity in 1909, was between tall at the shoulders, and had a thick, falling
mane
Mane may refer to:
* Mane (horse), the line of hair along the spine of the neck
* Mane (lion), the hair found around the male mammal's neck
In arts and entertainment
* ''Mane'' (film) is a 1990 Kannada language film directed by Girish Kasaravall ...
, a
grullo
Grulla or grullo, also called blue dun, gray dun or mouse dun, is a equine coat color, color of horses in the dun (color), dun family, characterized by tan-gray or mouse-colored hairs on the body, often with shoulder and dorsal stripes and black ...
coat colour, dark legs, and primitive markings, including a dorsal stripe and shoulder stripes.
A number of coat color
genotype
The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
s have been identified within European wild horses from the
Pleistocene
The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
and
Holocene
The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
: those creating
bay
A bay is a recessed, coastal body of water that directly connects to a larger main body of water, such as an ocean, a lake, or another bay. A large bay is usually called a Gulf (geography), gulf, sea, sound (geography), sound, or bight (geogra ...
,
black
Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white ha ...
and leopard complex are known from the wild horse population in Europe and were depicted in
cave painting
In archaeology, Cave paintings are a type of parietal art (which category also includes petroglyphs, or engravings), found on the wall or ceilings of caves. The term usually implies prehistoric origin, and the oldest known are more than 40,00 ...
s of wild horses during the Pleistocene. The
dun gene
The dun gene is a dilution gene that affects both red and black pigments in the equine coat color, coat color of a horse. The dun gene lightens most of the body while leaving the mane (horse), mane, tail, legs, and primitive markings the shade of ...
, a
dilution gene
A dilution gene is any one of a number of genes that act to create a lighter coat color in living creatures. There are many examples of such genes:
General
Diluted coat colors have melanocytes, but vary from darker colors due to the concentration ...
seen in
Przewalski's horse
Przewalski's horse (, , (Пржевальский ), ) (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered horse originally native to the steppes of Ce ...
that also creates the grullo or "blue dun" coat seen in the Konik, has not yet been studied in European wild horses. It is likely that at least some wild horses had a dun coat.
Whether or not the horses called tarpan had a standing mane like wild equines, or a falling mane like domestic horses cannot be ascertained anymore as historic reports have not been unambiguous on this subject.
The appearance of European wild horses may be reconstructed with genetic, osteologic and historic data. One genetic study suggests that bay was the predominant color in European wild horses. During the
Mesolithic
The Mesolithic (Greek: μέσος, ''mesos'' 'middle' + λίθος, ''lithos'' 'stone') or Middle Stone Age is the Old World archaeological period between the Upper Paleolithic and the Neolithic. The term Epipaleolithic is often used synonymo ...
, a gene coding a black coat color appeared on the Iberian peninsula. This color spread east but was less common than bay in the investigated sample and never reached Siberia. Bay in combination with dun results in the "bay dun" color seen in Przewalski's horses; black with dun creates the grullo coat. A loss of the dun dilution may have been advantageous in more forested western European landscapes, as dark colors were a better camouflage in forests.Baker, Sue, 2008: Exmoor Ponies: Survival of the Fittest – A natural history.Pangaré or "mealy" coloration, a characteristic of other wild equines, might have been present in at least some European wild horses, as historic accounts report a light belly.
Historic references describing the tarpan report that most of them were of a black dun (grullo, "blue dun") colour, while also black individuals and domestic colours like white legs or greys were reported. Because of that, some contemporaneous authors, such as Peter Pallas, believed tarpans to be escaped farm horses.Tadeusz Jezierski, Zbigniew Jaworski: Das Polnische Konik. Die Neue Brehm-Bücherei Bd. 658, Westarp Wissenschaften, Hohenwarsleben 2008
History
Wild horses have been present in Europe since the
Pleistocene
The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
and ranged from southern
France
France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan ar ...
and
Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond")
, national_anthem = (English: "Royal March")
, ...
east to central
Russia
Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
. There are cave drawings of primitive predomestication horses at
Lascaux
Lascaux ( , ; french: Grotte de Lascaux , "Lascaux Cave") is a network of caves near the village of Montignac, in the department of Dordogne in southwestern France. Over 600 parietal wall paintings cover the interior walls and ceilings of ...
, France and in Cave of Altamira, Spain, as well as artifacts believed to show the species in
southern Russia
Southern Russia or the South of Russia (russian: Юг России, ''Yug Rossii'') is a colloquial term for the southernmost geographic portion of European Russia generally covering the Southern Federal District and the North Caucasian Federal ...
, where a horse of this type was domesticated around 3000 BC. ''Equus ferus'' had a continuous range from western Europe to
Alaska
Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S ...
; historic material suggests wild horses lived in most parts of Holocene continental Europe and the Eurasian steppe, except for parts of Scandinavia, Iceland and Ireland.
Forest tarpan
The "forest horse" or "forest tarpan" was a hypothesis of various 20th-century natural scientists, including Tadeusz Vetulani, who suggested that the continuous forestation of Europe after the last ice age created forest-adapted subtype of the wild horse, which he named ''Equus sylvestris.'' However, historic references do not describe any major difference between the populations, and therefore most authors assume there was only one subspecies of western Eurasian wild horse, ''Equus ferus ferus.''
Nevertheless, a stocky type of horse living in forests and highlands was described during the 19th century in Spain, the
Pyrenees
The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
, the
Camargue
Camargue (, also , , ; oc, label=Provençal, Camarga) is a region of France located south of Arles, between the Mediterranean Sea and the two arms of the Rhône delta. The eastern arm is called the ''Grand Rhône''; the western one is the ...
, the
Ardennes
The Ardennes (french: Ardenne ; nl, Ardennen ; german: Ardennen; wa, Årdene ; lb, Ardennen ), also known as the Ardennes Forest or Forest of Ardennes, is a region of extensive forests, rough terrain, rolling hills and ridges primarily in Be ...
,
Great Britain
Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
, and the southern
Swedish
Swedish or ' may refer to:
Anything from or related to Sweden, a country in Northern Europe. Or, specifically:
* Swedish language, a North Germanic language spoken primarily in Sweden and Finland
** Swedish alphabet, the official alphabet used by ...
upland.
They had a robust head and strong body, and a long frizzy mane. The color was described as faint brown or yellowish brown with
eel stripe
Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are ...
and leg stripes, or wholly black legs. The flanks and shoulders were spotted, some of them tended to an ashy colour. They dwelled in rocky habitats and showed intelligent and fierce behaviour. Yet, those horses were never colloquially called "tarpans".Cis van Vuure: On the origin of the Polish konik and its relation to Dutch nature management. 2014
Black wild horses were found in Dutch swamps, with a large skull, small eyes, and a bristly muzzle. Their mane was full, with broad hooves, and curly hair. However, it is possible that these were feral and not wild horses.
Herodotus
Herodotus ( ; grc, , }; BC) was an ancient Greek historian and geographer from the Greek city of Halicarnassus, part of the Persian Empire (now Bodrum, Turkey) and a later citizen of Thurii in modern Calabria ( Italy). He is known for ...
described light-coloured wild horses in an area now part of
Ukraine
Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian invas ...
in the 5th century BCE. In the 12th century,
Albertus Magnus
Albertus Magnus (c. 1200 – 15 November 1280), also known as Saint Albert the Great or Albert of Cologne, was a German Dominican friar, philosopher, scientist, and bishop. Later canonised as a Catholic saint, he was known during his li ...
stated that mouse-coloured wild horses with a dark eel stripe lived in the German territory, and in Denmark, large herds were hunted.
16th century free-ranging horses in Europe
Wild horses still were common in the east of Prussia during the 15th and early 16th centuries. During the 16th century, wild horses disappeared from most of the mainland of western Europe and became less common in eastern Europe as well. Belsazar Hacquet saw wild horses in the Polish zoo in
Zamość
Zamość (; yi, זאמאשטש, Zamoshtsh; la, Zamoscia) is a historical city in southeastern Poland. It is situated in the southern part of Lublin Voivodeship, about from Lublin, from Warsaw. In 2021, the population of Zamość was 62,021.
...
during the
Seven Years' War
The Seven Years' War (1756–1763) was a global conflict that involved most of the European Great Powers, and was fought primarily in Europe, the Americas, and Asia-Pacific. Other concurrent conflicts include the French and Indian War (1754– ...
. According to him, those wild horses were of small body size, had a blackish brown colour, a large and thick head, short dark manes and tail hair, and a "beard". They were absolutely untameable and defended themselves harshly against predators.
Kajetan Kozmian visited the population at Zamość as well and reported that they were small and strong, had robust limbs and a consistently dark mouse colour. Samuel Gottlieb Gmelin witnessed herds in
Voronezh
Voronezh ( rus, links=no, Воро́неж, p=vɐˈronʲɪʂ}) is a city and the administrative centre of Voronezh Oblast in southwestern Russia straddling the Voronezh River, located from where it flows into the Don River. The city sits on ...
in 1768. Those horses were described as very fast and shy, fleeing at any noise, and as small, with small, pinned ears and a short frizzly mane. The tail was shorter than in domestic horses. They were typically mouse-colored with a light belly and legs becoming black, although
gray
Grey (more common in British English) or gray (more common in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning literally that it is "without color", because it can be composed ...
and white horses were mentioned as well. The coat was long and dense. The horses at Zamosc were never called "tarpan" back in their lifetime.
Peter Simon Pallas
Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810.
Life and work
Peter Simon Pallas was born in Berlin, the son ...
witnessed possible tarpans in the same year in southern Russia. He thought they were feral animals that escaped during the confusions of wars. These herds were important game of the Tatars and numbered between 5 and 20 animals. The horses he described had a small body, large and thick heads, short frizzly manes and short tail hair, as well as pinned ears. The colour was described as faint brownish, sometimes brown or black. He also reported of obvious domestic hybrids with light-colored legs or gray coats.
The tarpans of 18th century Europe
''The Natural History of Horses'' by 19th-century author Charles Hamilton Smith also described tarpans. According to Smith, the herds of free-ranging horses numbered from a few to hundred individuals. They often were mixed with domestic horses, and alongside pure herds there were herds of feral horses or hybrids. The color of alleged pure tarpans was described as consistently brown, cream-colored or mouse-colored. The short frizzy mane was reported to be black, as were the tail and legs. The ears were of varying size, but set high on the skull. The eyes were small.
According to Smith, tarpans made stronger sounds than domestic horses and the overall appearance of these horses was mule-like.
Extinction
The human-caused extinction of wild horses in Europe began in Southern Europe, possibly in antiquity. While humans had been hunting wild horses since the
Paleolithic
The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός '' palaios'', "old" and λίθος ''lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone to ...
, during historic times
horse meat
Horse meat forms a significant part of the culinary traditions of many countries, particularly in Eurasia. The eight countries that consume the most horse meat consume about 4.3 million horses a year. For the majority of humanity's early existen ...
was an important source of protein for many cultures. As large herbivores, the range of the tarpan was continuously decreased by the increasing human population of the Eurasian landmass. Wild horses were further targeted because they caused damage to hay stores and often took domestic mares from pastures. Furthermore, interbreeding with wild horses was an economic loss for farmers since the
foal
A foal is an equine up to one year old; this term is used mainly for horses, but can be used for donkeys. More specific terms are colt for a male foal and filly for a female foal, and are used until the horse is three or four. When the foal ...
s of such matings were intractable.
Tarpans lived in the southern parts of the Russian steppe. In 1879 the last scientifically confirmed individual was killed. After that, only dubious sightings were documented. As the tarpan horse died out in the wild between 1875 and 1890, the last free-ranging mare considered a tarpan was accidentally killed during an attempt at capture. The last captive tarpan died in 1909 in a Russian zoo.
The horses at Zamosc survived until 1806, when they were allegedly donated to local farmers of Poland and bred to their horses.Bunzel-Drüke, Finck, Kämmer, Luick, Reisinger, Riecken, Riedl, Scharf & Zimball: ''"Wilde Weiden: Praxisleitfaden für Ganzjahresbeweidung in Naturschutz und Landschaftsentwicklung''Thomas Jansen: ''Untersuchungen zur Phylogenie und Domestikation des Hauspferdes (Equus ferus f. caballus) Stammesentwicklung und geografische Verteilung.'' 2002 PDF ) The
Konik
The Konik or Polish Konik, pl, konik polski, is a Polish breed of pony. There are semi-feral populations in some regions. They are usually mouse dun or striped dun in color.
The Bilgoray, pl, konik biłgorajski, of south-eastern Poland is a ...
is claimed to descend from these hybrid horses, as recent research has highlighted a significant degree of anatomic difference between free-roaming Konik in the Netherlands and other modern domesticated horses.
Tarpans interbreeding with domestic horses
The oldest archaeological evidence for domesticated horses is from Kazakhstan and Ukraine between 6,000 and 5,500 YBP (years before present). The diverse mitochondrial DNA of domestic horses contrasts sharply with the very low diversity of the Y chromosome; that suggests that many mares but only a few stallions were used, and local use of wild mares or even secondary sites of domestication are likely. Therefore, the European wild horse may well have contributed to the domestic horse.
Wild horses vs. feral horses
Some researchers consider the tarpans to be mixed wild and feral population or completely feral horses. Few consider the more recent animals historically called "tarpans" to be genuine wild horses without domestic influence. Historic references to "wild horses" may actually refer to feral domestic horses or hybrids.
Some 19th century authors wrote that local "wild" horses had hoof problems that led to crippled legs; therefore, they assumed these were feral horses. Other contemporary authors claimed all "wild" horses between the Volga River and the Ural were actually feral. However, others thought that this was too speculative and assumed that wild, undomesticated horses still lived into the 19th century. Domestic horses used in warfare often were turned loose when they were no longer needed. Also, remaining wild stallions could steal domestic mares. There are some accounts from the 18th and 19th centuries of wild herds with typical wild horse features such as large heads, pinned ears, short frizzy mane and tail, but mentioned animals with domestic influence as well.
The only known individual to be photographed was the so-called Cherson tarpan, which was caught as a foal near
Novovorontsovka
Novovorontsovka ( uk, Нововоронцовка; russian: Нововоронцовка) is an urban-type settlement in Beryslav Raion, Kherson Oblast, southern Ukraine. It hosts the administration of Novovorontsovka settlement hromada, one of t ...
in 1866. It died in 1887 in the Moscow Zoo. The nature of this horse was dubious in its lifetime, because it showed almost none of the wild horse features described in the historic sources. In 2021, a study found that the so-called 'Shatilov' tarpan, a museum specimen from the
Kherson region
Kherson Oblast ( uk, Херсо́нська о́бласть, translit=Khersónsʹka óblastʹ, ), also known as Khersonshchyna ( uk, Херсо́нщина, ), is an oblast (province) in southern Ukraine, currently claimed and partly occupie ...
of the
Pontic–Caspian steppe
The Pontic–Caspian steppe, formed by the Caspian steppe and the Pontic steppe, is the steppeland stretching from the northern shores of the Black Sea (the Pontus Euxinus of antiquity) to the northern area around the Caspian Sea. It extends ...
that died in 1868, was a hybrid between two horse lineages, with two thirds of its genetics representing the same ancestral lineage as modern horses, and the remaining third related to a distinct lineage of European horses found at an archaeological site associated with the late-
Neolithic
The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several part ...
Corded Ware culture
The Corded Ware culture comprises a broad archaeological horizon of Europe between ca. 3000 BC – 2350 BC, thus from the late Neolithic, through the Copper Age, and ending in the early Bronze Age. Corded Ware culture encompassed a v ...
.
Breeding back the tarpan
Three attempts have been made to use
selective breeding
Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant ma ...
to create a type of horse that resembles the tarpan
phenotype
In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological prop ...
, though recreating an extinct subspecies is not genetically possible with current technology. In 1936, Polish university professor Tadeusz Vetulani selected Polish farm horses that were formerly known as Panje horses (now called Konik) and that he believed resembled the historic tarpan and started a
selective breeding
Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant ma ...
program. Although his breeding attempt is well known, it made only a minor contribution to the modern
Konik
The Konik or Polish Konik, pl, konik polski, is a Polish breed of pony. There are semi-feral populations in some regions. They are usually mouse dun or striped dun in color.
The Bilgoray, pl, konik biłgorajski, of south-eastern Poland is a ...
stock,Tadeusz Jezierski, Zbigniew Jaworski: ''Das Polnische Konik. Die Neue Brehm-Bücherei Bd. 658'', Westarp Wissenschaften, Hohenwarsleben 2008, which clusters genetically with other domestic horse breeds, including those as diverse as the Mongolian horse and the
Thoroughbred
The Thoroughbred is a horse breed best known for its use in horse racing. Although the word ''thoroughbred'' is sometimes used to refer to any breed of purebred horse, it technically refers only to the Thoroughbred breed. Thoroughbreds are ...
Munich Zoo
Hellabrunn Zoo (or Tierpark Hellabrunn in German) is a 40 hectare (99 acre) zoological garden in the Bavarian capital of Munich. The zoo is situated on the right bank of the river Isar, in the southern part of Munich near the quarter of Thalkirc ...
began a program crossbreeding Koniks with Przewalski horses, Gotland ponies, and Icelandic horses. By the 1960s they produced the Heck horse. In the mid-1960s, Harry Hegard started a similar program in the United States using mustangs and local working ranch horses that has resulted in the Hegardt or Stroebel's horse.
Assessment
While all three breeds have a primitive look that resembles the tarpan in some respects, they are not genetically tarpans and the wild, predomestic European horse remains extinct. However, this does not prevent some modern breeders from marketing horses with these features as a "tarpan".
In spite of sharing primitive external features, the Konik and Hucul horses have markedly different conformation with differently proportioned body measurements, thought in part to be linked to living in different habitats.
Other breeds sometimes alleged to be surviving wild horses include the
Exmoor pony
The Exmoor pony is one of the mountain and moorland pony breeds native to the British Isles. Some still roam as semi-feral livestock on Exmoor, a large area of moorland in Devon and Somerset in southwest England. The Exmoor has been give ...
and the Dülmen pony. However, genetic studies do not set any of these breeds apart from other domestic horses. On the other hand, there has not yet been a study comparing domestic breeds directly with the European wild horse.
See also
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Domestication of the horse
A number of hypotheses exist on many of the key issues regarding the domestication of the horse. Although horses appeared in Paleolithic cave art as early as 30,000 BCE, these were wild horses and were probably hunted for meat.
How and when ho ...
Przewalski's horse
Przewalski's horse (, , (Пржевальский ), ) (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered horse originally native to the steppes of Ce ...