Taproot Dieback
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A taproot is a large, central, and dominant
root In vascular plants, the roots are the plant organ, organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often bel ...
from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the
carrot The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in colour, though heirloom variants including purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild ...
, the taproot is a
storage organ A storage organ is a part of a plant specifically modified for storage of energy (generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...
so well developed that it has been cultivated as a vegetable. The taproot system contrasts with the adventitious- or
fibrous-root system A fibrous root system is the opposite of a taproot system. It is usually formed by thin, moderately branching roots growing from the stem. A fibrous root system is universal in monocotyledonous plants and ferns. The fibrous root systems look li ...
of plants with many branched roots, but many plants that grow a taproot during
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, ...
go on to develop branching root structures, although some that rely on the main root for storage may retain the dominant taproot for centuries—for example, ''
Welwitschia ''Welwitschia'' is a monotypic genus of gnetophytes containing only the species ''Welwitschia mirabilis''. It is named after the Austrian botanist Friedrich Welwitsch, who documented the plant in the 1850s. In common use, it is sometimes ref ...
''.


Description

Dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, ...
s, one of the two divisions of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s (angiosperms), start with a taproot, which is one main root forming from the enlarging
radicle In botany, the radicle is the first part of a seedling (a growing plant embryo) to emerge from the seed during the process of germination. The radicle is the embryonic root of the plant, and grows downward in the soil (the shoot emerges from t ...
of the seed. The tap root can be persistent throughout the life of the plant but is most often replaced later in the plant's development by a fibrous root system. A persistent taproot system forms when the radicle keeps growing and smaller lateral roots form along the taproot. The shape of taproots can vary but the typical shapes include: *Conical root: this type of root tuber is conical in shape, i.e. widest at the top and tapering steadily towards the bottom: e.g.
carrot The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in colour, though heirloom variants including purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild ...
. *Fusiform root: this root is widest in the middle and tapers towards the top and the bottom: e.g.
radish The radish (''Raphanus sativus'') is a flowering plant in the mustard family, Brassicaceae. Its large taproot is commonly used as a root vegetable, although the entire plant is edible and its leaves are sometimes used as a leaf vegetable. Origina ...
. *Napiform root: the root has a
top Top most commonly refers to: * Top, a basic term of orientation, distinguished from bottom, front, back, and sides * Spinning top, a ubiquitous traditional toy * Top (clothing), clothing designed to be worn over the torso * Mountain top, a moun ...
-like appearance. It is very broad at the top and tapers suddenly like a tail at the bottom: e.g.
turnip The turnip or white turnip ('' Brassica rapa'' subsp. ''rapa'') is a root vegetable commonly grown in temperate climates worldwide for its white, fleshy taproot. Small, tender varieties are grown for human consumption, while larger varieties a ...
. Many taproots are modified into
storage organ A storage organ is a part of a plant specifically modified for storage of energy (generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...
s. Some plants with taproots: *
Beetroot The beetroot (British English) or beet (North American English) is the taproot portion of a ''Beta vulgaris'' subsp. ''vulgaris'' plant in the Conditiva Group. The plant is a root vegetable also known as the table beet, garden beet, dinner bee ...
*
Burdock ''Arctium'' is a genus of biennial plants commonly known as burdock, family Asteraceae. Native to Europe and Asia, several species have been widely introduced worldwide. Burdock's clinging properties, in addition to providing an excellent mecha ...
*
Carrot The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in colour, though heirloom variants including purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild ...
*
Sugar beet A sugar beet is a plant whose root contains a high concentration of sucrose and that is grown commercially for sugar production. In plant breeding, it is known as the Altissima cultivar group of the common beet (''Beta vulgaris''). Together with ...
*
Dandelion ''Taraxacum'' () is a genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus has a near-cosmopolitan distribu ...
*
Parsley Parsley, or garden parsley (''Petroselinum crispum''), is a species of flowering plant in the family Apiaceae that is native to Greece, Morocco and the former Yugoslavia. It has been introduced and naturalisation (biology), naturalized in Eur ...
*
Parsnip The parsnip (''Pastinaca sativa'') is a root vegetable closely related to carrot and parsley, all belonging to the flowering plant family Apiaceae. It is a biennial plant usually grown as an annual. Its long taproot has cream-colored skin an ...
*
Poppy mallow ''Callirhoe'' is a genus of flowering plants in the Malva, mallow family, Malvaceae. Its nine species are commonly known as poppy mallows and all are native to the prairies and grasslands of North America. Of the nine species, some are Annual pla ...
*
Radish The radish (''Raphanus sativus'') is a flowering plant in the mustard family, Brassicaceae. Its large taproot is commonly used as a root vegetable, although the entire plant is edible and its leaves are sometimes used as a leaf vegetable. Origina ...
*
Sagebrush Sagebrush is the common name of several woody and herbaceous species of plants in the genus ''Artemisia (plant), Artemisia''. The best-known sagebrush is the shrub ''Artemisia tridentata''. Sagebrush is native to the western half of North Amer ...
*
Turnip The turnip or white turnip ('' Brassica rapa'' subsp. ''rapa'') is a root vegetable commonly grown in temperate climates worldwide for its white, fleshy taproot. Small, tender varieties are grown for human consumption, while larger varieties a ...
*
Common milkweed ''Asclepias syriaca'', commonly called common milkweed, butterfly flower, silkweed, silky swallow-wort, and Virginia silkweed, is a species of flowering plant. It is native to southern Canada and much of the United States east of the Rocky Moun ...
* trees such as
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
s,
elm Elms are deciduous and semi-deciduous trees comprising the genus ''Ulmus'' in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical- montane regions of North America and Eurasia, ...
s,
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
s and
fir Firs are evergreen coniferous trees belonging to the genus ''Abies'' () in the family Pinaceae. There are approximately 48–65 extant species, found on mountains throughout much of North and Central America, Eurasia, and North Africa. The genu ...
s


Development

Taproots develop from the
radicle In botany, the radicle is the first part of a seedling (a growing plant embryo) to emerge from the seed during the process of germination. The radicle is the embryonic root of the plant, and grows downward in the soil (the shoot emerges from t ...
of a seed, forming the primary root. It branches off to secondary roots, which in turn branch to form tertiary roots. These may further branch to form rootlets. For most plants species the radicle dies some time after seed germination, causing the development of a fibrous root system, which lacks a main downward-growing root. Most
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s begin life with a taproot, but after one to a few years the main root system changes to a wide-spreading fibrous root system with mainly horizontal-growing surface roots and only a few vertical, deep-anchoring roots. A typical mature tree 30–50 m tall has a root system that extends horizontally in all directions as far as the tree is tall or more, but as much as 100% of the roots are in the top 50 cm of soil. Soil characteristics strongly influence the architecture of taproots; for example, deep and rich soils favour the development of vertical taproots in many oak species such as ''
Quercus kelloggii ''Quercus kelloggii'', the California black oak or Kellogg oak, is an oak in the red oak section (genus ''Quercus'', section ''Lobatae'', series ''Agrifoliae'') native to western North America. Description ''Quercus kelloggii'' grows from one ...
'', while clay soils promote the growth of multiple taproots.


Horticultural considerations

Many plants with taproots are difficult to transplant, or even to grow in containers, because the root tends to grow deep rapidly and in many species comparatively slight obstacles or damage to the taproot will stunt or kill the plant. Among weeds with taproots
dandelion ''Taraxacum'' () is a genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus has a near-cosmopolitan distribu ...
s are typical; being deep-rooted, they are hard to uproot and if the taproot breaks off near the top, the part that stays in the ground often resprouts such that, for effective control, the taproot needs to be severed at least several centimetres below ground level.


Gallery

file:Buckeye6.jpg, A tree taproot file:Plant taproots.jpg, A taproot


References


External links

*
Fullerton Arboretum on taproots
{{Webarchive, url=https://web.archive.org/web/20080308052559/http://arboretum.fullerton.edu/grow/taproot.asp , date=2008-03-08 Plant roots