Tapirus Indicus
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The Malayan tapir (''Tapirus indicus''), also called Asian tapir, Asiatic tapir, oriental tapir, Indian tapir, piebald tapir, or black-and-white tapir, is the only living
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a Suidae, pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South America, South and Centr ...
species outside of the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
. It is native to
Southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
from the
Malay Peninsula The Malay Peninsula is located in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area contains Peninsular Malaysia, Southern Tha ...
to
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
. It has been listed as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
since 2008, as the population is estimated to comprise fewer than 2,500 mature individuals.


Taxonomy

The
scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
''Tapirus indicus'' was proposed by
Anselme Gaëtan Desmarest Anselme Gaëtan Desmarest (6 March 1784 – 4 June 1838) was a French Zoology, zoologist and author. He was the son of Nicolas Desmarest and the father of Eugène Anselme Sébastien Léon Desmarest. Career Desmarest was a disciple of Georges Cu ...
in 1819 who referred to a tapir described by
Pierre-Médard Diard Pierre-Médard Diard (; 19 March 1794 – 16 February 1863) was a French natural history, naturalist and explorer. Diard studied zoology and anatomy under Georges Cuvier and assisted him in researches on the development of the foetus and on the eg ...
. ''Tapirus indicus brevetianus'' was coined by a Dutch zoologist in 1926 who described a black Malayan tapir from
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
that had been sent to
Rotterdam Zoo (; ), officially Rotterdam Zoo, is a zoo located in the northwestern part of Rotterdam. It is one of the oldest zoos in the Netherlands, and has been operated by the ("Royal Rotterdam Zoo Foundation"). Divided into several zoogeographic regio ...
in the early 1920s.
Phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analyses of 13 Malayan tapirs showed that the species is
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
. It was placed in the genus ''Acrocodia'' by
Colin Groves Colin Peter Groves (24 June 1942 – 30 November 2017) was a British-Australian biologist and anthropologist. Groves was professor of biological anthropology at the Australian National University in Canberra, Australia. Education Born in Englan ...
and Peter Grubb in 2011. However, a comparison of
mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
of 16
perissodactyl Perissodactyla (, ), or odd-toed ungulates, is an order of Ungulate, ungulates. The order includes about 17 living species divided into three Family (biology), families: Equidae (wild horse, horses, Asinus, asses, and zebras), Rhinocerotidae ( ...
species revealed that the Malayan tapir forms a
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
together with the ''
Tapirus ''Tapirus'' is a genus of tapir which contains the living tapir species. The Malayan tapir is usually included in ''Tapirus'' as well, although some authorities have moved it into its own genus, ''Acrocodia''. Extant species The Kabomani tapir ...
'' species native to the Americas. It was the first ''Tapirus'' species that genetically diverged from the group, estimated about in the
Late Oligocene The Chattian is, in the geologic timescale The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the pro ...
.


Description

The Malayan tapir is easily identified by its markings, most notably the light-colored patch that extends from its shoulders to its hindquarters. Black hair covers its head, shoulders, and legs, while white hair covers its midsection, rear, and the tips of its ears; these white edges around the rims of the outer ear as is true of other tapirs. The disrupted coloration breaks up its outline, providing
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
by making the animal difficult to recognize against the varied terrain and dense flora of its habitat; potential
predators Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
may mistake it for a large rock, rather than
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
, when it is lying down to sleep. The Malayan tapir is the largest of the four extant tapir species and grows to between in length, not counting a stubby tail of only in length, and stands tall. It typically weighs between , although some adults can weigh up to .Tapirus indicus
Animal Diversity Web
Asian Tapir
,, Arkive
The females are usually larger than the males. Like other tapir species, it has a small, stubby tail and a long, flexible
proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular arthropod mouthparts, mouthparts used for feeding and sucking. In vertebrates, a pr ...
. It has four toes on each front foot and three toes on each back foot. The Malayan tapir has rather poor eyesight, but excellent hearing and sense of smell. The tapir's unique proboscis is supported by several evolutionary adaptations of its
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
. It has a large
sagittal crest A sagittal crest is a ridge of bone running lengthwise along the midline of the top of the skull (at the sagittal suture) of many mammalian and reptilian skulls, among others. The presence of this ridge of bone indicates that there are excepti ...
, unusually positioned orbits, an unusually shaped cranium with elevated frontal bones, and a retracted nasal incision as well as retracted facial cartilage. This evolutionary process is believed to have caused the loss of some cartilages, facial muscles, and the bony wall of the tapir's nasal chamber.


Vision

Malayan tapirs have very poor eyesight, both on land and in water, instead relying heavily on their excellent senses of smell and hearing to navigate and forage. Their eyes are small and, like many herbivores, positioned on the sides of the face. They have brown irises, but the
corneas The cornea is the transparency (optics), transparent front part of the eyeball which covers the Iris (anatomy), iris, pupil, and Anterior chamber of eyeball, anterior chamber. Along with the anterior chamber and Lens (anatomy), lens, the cornea ...
are often covered in a blue haze; this corneal cloudiness is thought to be caused by repetitive exposure to light. This loss of transparency impacts the ability of the cornea to transmit and focus outside light as it enters the eye, impairing the animal's overall vision. As these tapirs are most active at night on top of having poor eyesight, this habit may make it harder for them to search for food and avoid predators.


Color variation

Two melanistic Malayan tapirs were observed in Jerangau Forest Reserve in Malaysia in 2000. A black Malayan tapir was also recorded in Tekai Tembeling Forest Reserve in
Pahang {{Infobox political division , name = Pahang , official_name = Pahang Darul Makmur , native_name = , settlement_type = States and federal territories of Malaysia, State , image_skyline = , imagesize ...
state in 2016.


Distribution and habitat

The Malayan tapir lives throughout primary and secondary tropical lowland
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s of
Southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
, including
Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
in
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
,
Peninsular Malaysia Peninsular Malaysia, historically known as Malaya and also known as West Malaysia or the Malaysian Peninsula, is the western part of Malaysia that comprises the southern part of the Malay Peninsula on Mainland Southeast Asia and the list of isla ...
,
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
, and
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
.
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s were found in
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...
and other locations accompanied by herbivores more typical of grasslands, indicating that it evolved in more open habitats and retreated to closed forests in later times. It was found in
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
until at least 8,000 years ago during the early
Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
in the
Niah Caves Niah National Park, located within Miri Division, Sarawak, Malaysia, is the site of the Niah Caves which are an archeological site. History Alfred Russel Wallace lived for 8 months at Simunjan District with a mining engineer, Robert Coulson, ...
of
Sarawak Sarawak ( , ) is a States and federal territories of Malaysia, state of Malaysia. It is the largest among the 13 states, with an area almost equal to that of Peninsular Malaysia. Sarawak is located in East Malaysia in northwest Borneo, and is ...
, and some 19th century writers mentioned it as a contemporary species in Borneo, likely based on native accounts. It has been proposed to reintroduce the tapir to the island as a conservation measure. In the continent, the Malayan tapir was found in historical times as far north as
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
.


Behaviour and ecology

Malayan tapirs are primarily solitary, marking out large tracts of land as their
territory A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
, though these areas usually overlap with those of other individuals. Tapirs mark out their territories by
spraying urine Urination is the release of urine from the bladder through the urethra in placental mammals, or through the cloaca in other vertebrates. It is the urinary system's form of excretion. It is also known medically as micturition, voiding, ure ...
on plants, and they often follow distinct paths which they have bulldozed through the undergrowth. Exclusively
herbivorous A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
, the animal forages for the tender shoots and leaves of more than 115 species of plants, of which around 30 are particularly preferred, moving slowly through the
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
and pausing often to eat and note the scents left behind by other tapirs in the area. The tapir can run quickly when threatened or frightened, and if forced to fight can defend itself with its strong jaws and sharp teeth. Malayan tapirs communicate with high-pitched squeaks and whistles. They usually prefer to live near water and often bathe and swim, and they are also able to climb steep slopes. Tapirs are mainly active at night, though they are not exclusively
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
; because they tend to eat soon after sunset or before sunrise, and they will often nap in the middle of the night, they are considered to be
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal (active during dawn), vespertine (biology), vespertine/vespertinal (active during dusk), or both. This is distinguished from diurnalit ...
animals.


Life cycle

The
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...
period of the Malayan tapir is about 390–395 days, after which a single calf is born that weighs around . Malayan tapirs are the largest of the four tapir species at birth and tend to grow more quickly than their relatives. Young tapirs of all species have brown hair with white stripes and spots, a pattern that enables them to hide effectively in the dappled light of the forest. This baby coat fades into adult coloration between four and seven months after birth.
Weaning Weaning is the process of gradually introducing an infant human or other mammal to what will be its adult diet while withdrawing the supply of its mother's milk. In the United Kingdom, UK, weaning primarily refers to the introduction of solid ...
occurs between six and eight months of age, at which time the babies are nearly full-grown, and the animals reach
sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...
around age three.
Breeding Breeding is sexual reproduction that produces offspring, usually animals or plants. It can only occur between a male and a female animal or plant. Breeding may refer to: * Animal husbandry, through selected specimens such as dogs, horses, and rab ...
typically occurs in April, May or June, and females generally produce one calf every two years. Malayan tapirs can live up to 30 years, both in the wild and in captivity.


Predators

Because of its size, the Malayan tapir has few natural predators, and even reports of killings by
tiger The tiger (''Panthera tigris'') is a large Felidae, cat and a member of the genus ''Panthera'' native to Asia. It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly vertical stripes. It is ...
s (''Panthera tigris'') are scarce. Malayan tapirs can defend themselves with their very powerful bite; in 1998, the bite of a captive female Malayan tapir severed off a zookeeper's left arm at the mid-bicep, likely because she stood between her and her offspring.


Threats

The main threats to the Malayan tapir are loss and destruction of habitat through
deforestation Deforestation or forest clearance is the removal and destruction of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. Ab ...
. Large tracts of forests in Thailand and Malaysia have been converted for planting
oil palms ''Elaeis'' () is a genus of palms, called oil palms, containing two species, native to Africa and the Americas. They are used in commercial agriculture in the production of palm oil. Description Mature palms are single-stemmed, and can grow ...
. Habitat fragmentation in peninsular Malaysia caused displacement of 142 Malayan tapirs between 2006 and 2010; some were rescued and relocated, while 15 of them were killed in vehicle collisions.


References


External links

* *ARKive â€
images and movies of the Asian tapir ''(Tapirus indicus)''
*Tapir Specialist Group â€

{{Taxonbar, from1=Q192710, from2=Q27868280, from3=Q27868283 Tapirs
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a Suidae, pig, with a short, prehensile nose trunk (proboscis). Tapirs inhabit jungle and forest regions of South America, South and Centr ...
Mammals of Malaysia Mammals of Indonesia Mammals of Myanmar Mammals of Thailand Endangered fauna of Asia EDGE species Mammals described in 1819 Taxobox binomials not recognized by IUCN