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The tapaculos or tapacolos are a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
, Rhinocryptidae, of small suboscine
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
birds, found mainly in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
and with the highest diversity in the
Andean The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long and wide (widest between 18°S ...
regions. Three species ( Chocó, Tacarcuna, and the silvery-fronted) are found in southern
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
.


Description

Tapaculos are small to medium-sized birds, with a total length ranging from 10–24 cm (4–9½ in). These are terrestrial species that fly only poorly on their short wings. They have strong legs, well-suited to their habitat of grassland or forest undergrowth. The tail is cocked and pointed towards the head, and the name ''tapaculo'' probably derives from
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many countries in the Americas **Spanish cuisine **Spanish history **Spanish culture ...
for loincloth. Another possible explanation is that it originates from the
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
an name for the white-throated tapaculo, simply ''tapaculo'', which is an
onomatopoeic Onomatopoeia (or rarely echoism) is a type of word, or the process of creating a word, that phonetically imitates, resembles, or suggests the sound that it describes. Common onomatopoeias in English include animal noises such as ''oink'', '' ...
reference to its commonly heard song. While the majority of the family are small blackish or brown birds there are some larger and more colourful species. All tapaculos are skulking birds that frequently stay low in dense vegetation, even the larger, colorful species, and this renders them difficult to see. They are best located and – in the case of ''Scytalopus'' spp. – identified by their vocalisations. They feed on insects, seeds and other soft plant material with their pointed bills, and will scratch on the ground like a
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera's native range is restricted to Eura ...
. Most species lay two or three white eggs in a covered location, whether it be a burrow, a hole in a tree, or a domed nest.


Status and conservation

Some species have highly localized distributions and, being poor fliers, they easily become isolated in small populations.
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
currently (2007) consider one species vulnerable (''Scytalopus panamensis''), three species
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
( ''S. iraiensis'', '' S. rodriguezi'' and '' S. robbinsi''), and two species
critically endangered An IUCN Red List critically endangered (CR or sometimes CE) species is one that has been categorized by the International Union for Conservation of Nature as facing an extremely high risk of extinction in the wild. As of December 2023, of t ...
( ''Eleoscytalopus psychopompus'' and '' Merulaxis stresemanni''). The two critically endangered species are restricted to Atlantic forest of eastern Brazil, and were only recently rediscovered after several years without any records.


Taxonomy

The tapaculos were traditionally placed in a distinct
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Rhinocryptidae; more recent research indicates that according to analysis of mt and
nDNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. It ...
sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is cal ...
data, the tapaculos might be better merged into the Formicariidae as
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Rhinocryptini, as they are closer to the antthrushes than either is to the true antpittas. An alternative family name Pteroptochidae, has been used historically. The phylogenetic tree shown below is based on a large-scale genetic study of the
suboscines The Tyranni (suboscines) are a suborder of passerine birds that includes more than 1,000 species, a large majority of which are South American. It is named after the type genus '' Tyrannus''. These have a different anatomy of the syrinx musculat ...
by Michael Harvey and collaborators that was published in 2020. A high resolution version of the phylogenetic tree in Figure 1 is available from the first author's websit
here
The species numbers are from the list maintained by the
International Ornithologists' Union The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", ...
(IOC). The species-limits within the genus '' Scytalopus'' have historically been difficult to determine. The birds are highly cryptic, and identification using visual features often is impossible. Vocal and
biochemical Biochemistry, or biological chemistry, is the study of chemical processes within and relating to living organisms. A sub-discipline of both chemistry and biology, biochemistry may be divided into three fields: structural biology, enzymology, ...
data is typically needed to clarify the taxonomic status of the various populations. Several new species have been described in recent years (e.g. ''S. whitneyi'' and ''S. frankeae'' from
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
). The Brazilian taxa are similarly complex with several recently described species and considerable confusion surrounding the use of the scientific name '' Scytalopus speluncae''. Additionally, still undescribed species are known to exist, while some species as currently defined actually may include several species (e.g. the southern population of the large-footed tapaculo may represent a yet undescribed species). The confusing situation is perhaps best illustrated by the fact that only ten species were recognized in this genus in 1970, while the figure now is more than four times as high.


References


Cited sources

*


External links


Tapaculo videos
on the Internet Bird Collection {{Taxonbar, from=Q390189 Formicariidae * Taxa named by Alexander Wetmore