''Tantallognathus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of four-limbed

vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...

(

tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...

in a broad sense) from the Mississippian of

Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...

. It is based on a small jaw fragment which shows similarities to ''

Crassigyrinus ''Crassigyrinus'' (from , 'thick' and , 'tadpole') is an extinct genus of carnivorous stem tetrapod from the Early Carboniferous Clackmannan Group of Scotland and possibly Greer, West Virginia. Discovery The type specimen was originally descri ...

'', baphetids, and

crown group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor ...

tetrapods. This fossil was found near

Tantallon Castle Tantallon Castle is a ruined mid-14th-century fortress, located east of North Berwick, in East Lothian, Scotland. It sits atop a promontory opposite the Bass Rock, looking out onto the Firth of Forth. The last medieval curtain wall castle to b ...

(namesake of the genus), and the species name honors famed Scottish fossil collector Stan Wood. ''Tantallognathus'' is one of the most advanced tetrapods found in the

Ballagan Formation The Ballagan Formation is a Formation (geology), geologic formation in Scotland and England. It preserves fossils dating back to the early part of the Carboniferous Period (geology), period (Tournaisian – early Visean). Its name comes from the ...

, a geological unit known for a diverse fauna of

stegocephalia Stegocephali (often spelled Stegocephalia, from Greek , lit. "roofed head") is a clade of vertebrate animals containing all fully limbed tetrapodomorphs. It is equivalent to a broad definition of the superclass Tetrapoda: under this broad ...

ns. Like other Ballagan Formation vertebrates, it helps to clarify a pulse of tetrapod evolution during

Romer's gap Romer's gap is an apparent gap in the Paleozoic tetrapod fossil record noted in the studies of paleontology and evolutionary biology, which represent periods in the Early Carboniferous from which excavators have not yet found relevant transition ...

, a time interval when fossils of tetrapods and their relatives are otherwise very rare.

References

Stegocephali Tournaisian life Viséan life Mississippian sarcopterygians of Europe Fossil taxa described in 2018 {{Paleo-tetrapodomorph-stub