Taldycupedidae is an extinct family of beetles primarily known from the

Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...

period.

Taxonomy

Kirejtshuk (2020) included the following genera in Taldycupedidae, preliminarily treating it as a subfamily Taldycupedinae of the family

Permocupedidae Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the Middle Permian. They became rare in the Late Permian, with only one specie ...

. * '' Afrotaldycupes'' –

Beaufort Group The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably under ...

, South Africa, Late Permian (Changhsingian) ** ''Afrotaldycupes africanus'' (Type species) ** ''Afrotaldycupes lidgettoniensis'' * '' Mesothoris'' –

Blackstone Formation, Australia The Blackstone Formation is a geologic formation of the Ipswich Coal Measures Group in southeastern Queensland, Australia, dating to the Carnian to Norian stages of the Late Triassic. The shales, siltstones, coal and tuffs were deposited in a la ...

Late Triassic (Norian) ** ''Mesothoris clathrata'' (Type species) ** ''Mesothoris grandis'' ** ''Mesothoris punctomarginum'' * '' Proterocupes'' (syn ''Lobanovia'' ) ** ''Proterocupes major'' –

Vokhma Formation Vokhma () is a rural locality (a ''settlement'') and the administrative center of Vokhomsky District, Kostroma Oblast, Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countrie ...

Vologda Vologda (, ) is a types of inhabited localities in Russia, city and the administrative center of Vologda Oblast, Russia, located on the river Vologda (river), Vologda within the watershed of the Northern Dvina. Population: The city serves as ...

, Russia, Late Permian (Changhsingian) ** ''Proterocupes nedubrovensis'' – Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian) ** ''Proterocupes permiana'' (Type species) – Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian) * '' Simmondsia'' – Blackstone Formation, Australia Late Triassic (Norian) ** ''Simmondsia cylindrica'' ** ''Simmondsia subpiriformis'' (Type species) * '' Taldycupes'' (syn ''Cryptocupes'' , ''Taldycupidium'' ) ** ''Taldycupes bergi'' – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian ** ''Taldycupes cellulosus'' – Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian) ** ''Taldycupes khalfini'' (Type species) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian ** ''Taldycupes moltshanovi'' – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian ** ''Taldycupes pingi'' –

Yinping Formation Yicheng District () is a district of Zaozhuang, Shandong, China. It has an area of and around 360,200 inhabitants (2003). As of the end of 2022, the total household population of Yicheng District was 417,086,000 people. Administrative divisions ...

, Anhui, China, Middle Permian (Capitanian) ** ''Taldycupes reticulatus'' – Akkolka Formation, Kazakhstan, Late Permian (Changhsingian) ** ''Taldycupes rosanovi'' – Limptekon Formation,

Krasnoyarsk Krasnoyarsk is the largest types of inhabited localities in Russia, city and administrative center of Krasnoyarsk Krai, Russia. It is situated along the Yenisey, Yenisey River, and is the second-largest city in Siberia after Novosibirsk, with a p ...

, Russia, Late Permian (Changhsingian) ** ''Taldycupes vasjuchitshevi'' – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian * '' Tecticupes'' (syn ''Stegocupes'' ) ** ''Tecticupes heckeri'' (Type species) – Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian) Mitina Formation, Kemerovo, Russia, Middle Permian (Roadian) ** ''Tecticupes indistinctus'' – Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian) ** ''Tecticupes martynovi'' – Amanak Formation. Orenburg, Middle Permian (Capitanian) * '' Tychticupes'' – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian ** ''Tychticupes beljanini'' ** ''Tychticupes radtshenkoi'' (Type species) ** ''Tychticupes ragozini'' ** ''"Tychticupes" ussovi'' (

nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...

) * '' Tychticupoides'' – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian ** ''Tychticupoides grjazevi'' (Type species) * '' Uskaticupes'' (

) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian ** ''Uskaticupes javorskyi'' (

) The genera '' Alveolacupes'' , '' Clathropenna'' and '' Penecupes'' were previously placed in the family by some authors, but were all excluded from it by Kirejtshuk (2020). The genera '' Wuchangia'' and '' Yuxianocoleus'' , both from the Lower Jurassic of Hubei, China, were also excluded from the family by Zhao ''et al.'' (2021) (as the subfamily Taldycupedinae).

References

{{Taxonbar, from=Q18615698 Permocupedoidea Permian insects Taxa named by Boris Rohdendorf

† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species or languages). It is one of the mo ...

Prehistoric insect families