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Armen Leonovich Takhtajan or Takhtajian (; surname also transliterated Takhtadjan, Takhtadzhi︠a︡n or Takhtadzhian, pronounced takh-tuh-JAHN; 10 June 1910 – 13 November 2009), was a
Soviet The Union of Soviet Socialist Republics. (USSR), commonly known as the Soviet Union, was a List of former transcontinental countries#Since 1700, transcontinental country that spanned much of Eurasia from 1922 until Dissolution of the Soviet ...
-
Armenian Armenian may refer to: * Something of, from, or related to Armenia, a country in the South Caucasus region of Eurasia * Armenians, the national people of Armenia, or people of Armenian descent ** Armenian diaspora, Armenian communities around the ...
botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
, one of the most important figures in 20th century
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
and
systematics Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phy ...
and
biogeography Biogeography is the study of the species distribution, distribution of species and ecosystems in geography, geographic space and through evolutionary history of life, geological time. Organisms and biological community (ecology), communities o ...
. His other interests included
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s,
paleobotany Paleobotany or palaeobotany, also known as paleophytology, is the branch of botany dealing with the recovery and identification of plant fossils from geological contexts, and their use for the biological reconstruction of past environments ( pal ...
, and the flora of the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
. He was one of the most influential taxonomists of the latter twentieth century.


Life


Family

Takhtajan was born in
Shushi Shusha (, ) or Shushi () is a city in Azerbaijan, in the region of Nagorno-Karabakh. Situated at an altitude of 1,400–1,800 metres (4,600–5,900 ft) in the Karabakh mountains, the city was a mountain resort in the Soviet Union, Soviet ...
, Russian Empire, present-day disputed Nagorno-Karabakh, on 10 June 1910, to a family of
Armenian Armenian may refer to: * Something of, from, or related to Armenia, a country in the South Caucasus region of Eurasia * Armenians, the national people of Armenia, or people of Armenian descent ** Armenian diaspora, Armenian communities around the ...
intellectuals. His grandfather Meliksan Takhtadzhyan Petrovich had been born in
Trabzon Trabzon, historically known as Trebizond, is a city on the Black Sea coast of northeastern Turkey and the capital of Trabzon Province. The city was founded in 756 BC as "Trapezous" by colonists from Miletus. It was added into the Achaemenid E ...
, Ottoman Empire and was educated in Italy, on the island of San Lazzaro degli Armeni, an Armenian
enclave An enclave is a territory that is entirely surrounded by the territory of only one other state or entity. An enclave can be an independent territory or part of a larger one. Enclaves may also exist within territorial waters. ''Enclave'' is so ...
, spoke many languages and worked as a journalist. He died in Paris in 1930. His father, Leon Meliksanovich Takhtadzhyan (1884–1950), was born in
Batumi Batumi (; ka, ბათუმი ), historically Batum or Batoum, is the List of cities and towns in Georgia (country), second-largest city of Georgia (country), Georgia and the capital of the Autonomous Republic of Adjara, located on the coast ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
and was educated as an
agronomist An agriculturist, agriculturalist, agrologist, or agronomist (abbreviated as agr.) is a professional in the science, practice, and management of agriculture and agribusiness. It is a regulated profession in Canada, India, the Philippines, the Uni ...
at
Leipzig University Leipzig University (), in Leipzig in Saxony, Germany, is one of the world's oldest universities and the second-oldest university (by consecutive years of existence) in Germany. The university was founded on 2 December 1409 by Frederick I, Electo ...
. Graduating in 1906, he worked on farms in France, Switzerland and the United Kingdom, and made a special study of sheep farming. He became proficient in
German German(s) may refer to: * Germany, the country of the Germans and German things **Germania (Roman era) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizenship in Germany, see also Ge ...
, French, English,
Russian Russian(s) may refer to: *Russians (), an ethnic group of the East Slavic peoples, primarily living in Russia and neighboring countries *A citizen of Russia *Russian language, the most widely spoken of the Slavic languages *''The Russians'', a b ...
, Georgian and Azerbaijani. Arriving in Shushi in 1908, then a centre of sheep farming in the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
, looking for work, Leon was forced to teach German at the local ''
Realschule Real school (, ) is a type of secondary school in Germany, Switzerland and Liechtenstein. It has also existed in Croatia (''realna gimnazija''), the Austrian Empire, the German Empire, Denmark and Norway (''realskole''), Sweden (''realskola''), F ...
'' attached to the Armenian
seminary A seminary, school of theology, theological college, or divinity school is an educational institution for educating students (sometimes called seminarians) in scripture and theology, generally to prepare them for ordination to serve as cle ...
, due to lack of opportunities in his chosen field. There he met and married Gerseliya Sergeevna Gazarbekyan (1887–1974), Armen Takhtajan's mother, a native of Susha, in 1909.


Early life and education

The Takhtajans had three children, Armen (1910–2009), Nellie (1914–1994) and Nora (1918–1965). In 1918 the family were forced to flee to northern Armenia because of the
pogroms A pogrom is a violent riot incited with the aim of massacring or expelling an ethnic or religious group, particularly Jews. The term entered the English language from Russian to describe late 19th- and early 20th-century attacks on Jews i ...
. Throughout his childhood, Armen showed a keen interest in natural history, travelling with his father. Armen attended school in
Tbilisi Tbilisi ( ; ka, თბილისი, ), in some languages still known by its pre-1936 name Tiflis ( ), ( ka, ტფილისი, tr ) is the Capital city, capital and List of cities and towns in Georgia (country), largest city of Georgia ( ...
in nearby Georgia, initially at Unified Labor School number 42 (former Mantashevskom commercial school). There he came under the influence of one of his teachers, Alexander Konstantinovich Makaev (Makashvili) (1896–1962), who had previously taught agriculture at
Tbilisi State University Ivane Javakhishvili Tbilisi State University ( ka, ივანე ჯავახიშვილის სახელობის თბილისის სახელმწიფო უნივერსიტეტი, tr; often shorten ...
, and had produced a dictionary of botanical names in Georgian, Russian and Latin. Makaev would take Armen on botanical excursions, teaching him to identify plants from Sosnowski and Grossheim's "Determinants of plant life in the vicinity of Tbilisi" (1920). In 1928 he completed secondary school and travelled to
Leningrad Saint Petersburg, formerly known as Petrograd and later Leningrad, is the List of cities and towns in Russia by population, second-largest city in Russia after Moscow. It is situated on the Neva, River Neva, at the head of the Gulf of Finland ...
. There he volunteered at the biology school at
Leningrad University Saint Petersburg State University (SPBGU; ) is a public university, public research university in Saint Petersburg, Russia, and one of the oldest and most prestigious universities in Russia. Founded in 1724 by a decree of Peter the Great, the uni ...
and attended lectures by
Vladimir Leontyevich Komarov Vladimir Leontyevich Komarov (; – 5 December 1945) was a Russian and Soviet botanist. Biography Komarov was born in 1869. He was a graduate of Saint Petersburg State University, St. Petersburg University where he received a degree in bo ...
(1869–1945) on
plant morphology Phytomorphology is the study of the physical form and external structure of plants.Raven, P. H., R. F. Evert, & S. E. Eichhorn. ''Biology of Plants'', 7th ed., page 9. (New York: W. H. Freeman, 2005). . This is usually considered distinct from pl ...
. In 1929 he began his studies in biology at
Yerevan State University Yerevan State University (YSU; , , ), also simply University of Yerevan, is the oldest continuously operating public university in Armenia. Founded in 1919, it is the largest university in the country. It is thus informally known as Armenia's ...
in
Yerevan Yerevan ( , , ; ; sometimes spelled Erevan) is the capital and largest city of Armenia, as well as one of the world's List of oldest continuously inhabited cities, oldest continuously inhabited cities. Situated along the Hrazdan River, Yerev ...
, Armenia, which he completed in 1931. He then returned to Tbilisi, enrolling in the All-Union Institute of Subtropical Crops. In 1932 after completing his course at Tbilisi he worked for a while as a laboratory assistant at
Sukhumi Sukhumi or Sokhumi is a city in a wide bay on the Black Sea's eastern coast. It is both the Capital city, capital and largest city of Abkhazia, a partially recognised state that most countries consider a part of Georgia (country), Georgia. The ...
, Georgia, at the subtropical branch of the All-Union Institute of Applied Botany and New Crops (now the Institute of Plant Industry), before returning to Yerevan. In Yerevan he took a position as researcher at the Natural History Museum of Armenia, and then at the Herbarium of the Armenian branch of the Institute of Biology, Soviet Academy of Sciences, and began teaching at Yerevan University in 1936, while completing his Master's thesis. He died in
Saint Petersburg Saint Petersburg, formerly known as Petrograd and later Leningrad, is the List of cities and towns in Russia by population, second-largest city in Russia after Moscow. It is situated on the Neva, River Neva, at the head of the Gulf of Finland ...
on 13 November 2009, at the age of 99, in 2009, having just completed his most important work, ''Flowering Plants''.


Work

From 1938 to 1948 he headed a Department at the
Yerevan State University Yerevan State University (YSU; , , ), also simply University of Yerevan, is the oldest continuously operating public university in Armenia. Founded in 1919, it is the largest university in the country. It is thus informally known as Armenia's ...
, and in 1944–1948 he was director of the Botanical Institute of the Academy of Sciences of the Armenian SSR, and Professor of the Leningrad State University. Takhtajan was a member of the
Russian Academy of Sciences The Russian Academy of Sciences (RAS; ''Rossíyskaya akadémiya naúk'') consists of the national academy of Russia; a network of scientific research institutes from across the Russian Federation; and additional scientific and social units such ...
, as well as a foreign associate of the U.S.
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, NGO, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the ...
since 1971. He was also the academician of the Academy of Sciences of the Armenian SSR, the president of the Soviet All-Union Botanical Society (1973) and the International Association for Plant Taxonomy (1975), member of the Finnish Academy of Science and Literature (1971), the German Academy of Naturalists "Leopoldina" (1972) and other scientific societies. He was a member of the
Norwegian Academy of Science and Letters The Norwegian Academy of Science and Letters (, DNVA) is a learned society based in Oslo, Norway. Its purpose is to support the advancement of science and scholarship in Norway. History The Royal Frederick University in Christiania was establis ...
from 1980. While at the
Komarov Botanical Institute The Komarov Botanical Institute of the Russian Academy of Sciences () is a leading botanical institution in Russia, It is located on Aptekarsky Island in St. Petersburg, and is named after the Russian botanist Vladimir Leontyevich Komarov (1869– ...
in
Leningrad Saint Petersburg, formerly known as Petrograd and later Leningrad, is the List of cities and towns in Russia by population, second-largest city in Russia after Moscow. It is situated on the Neva, River Neva, at the head of the Gulf of Finland ...
in 1940, Takhtajan developed his classification scheme for
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s, which emphasized
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
relationships between plants. His system did not become known to botanists in the
West West is one of the four cardinal directions or points of the compass. It is the opposite direction from east and is the direction in which the Sun sets on the Earth. Etymology The word "west" is a Germanic word passed into some Romance langu ...
until after 1950, and in the late 1950s he began a correspondence and collaboration with the prominent American botanist Arthur Cronquist, whose plant classification scheme was heavily influenced by his collaboration with Takhtajan and other botanists at Komarov. He is chiefly famous as the author of works on the origins of flowering plants and paleobotany, developing a new classification system of higher plants. He worked on the "Flora of Armenia" (vol. 1–6, 1954–73) and "Fossil flowering plants of the USSR "(v. 1, 1974) books. Takhtajan also developed a system of
floristic region A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both re ...
s.(Takhtajan, Crovello and Cronquist, 1986) For many years restrictions were placed on his work because of his opposition to the official line on genetics promoted by Lysenko. In 1993 he worked for a while at the
New York Botanical Garden The New York Botanical Garden (NYBG) is a botanical garden at Bronx Park in the Bronx, New York City. Established in 1891, it is located on a site that contains a landscape with over one million living plants; the Enid A. Haupt Conservatory, ...
.


Takhtajan system

The "
Takhtajan system A list of systems of plant taxonomy, system of plant taxonomy, the Takhtajan system of plant classification was published by Armen Takhtajan, in several versions from the 1950s onwards. It is usually compared to the Cronquist system. It admits Para ...
" of flowering plant classification treats flowering plants as a division (phylum), '' Magnoliophyta'', with two classes, '' Magnoliopsida'' (dicots) and '' Liliopsida'' (monocots). These two classes are subdivided into subclasses, and then superorders, orders, and families. The Takhtajan system is similar to the
Cronquist system The Cronquist system is a list of systems of plant taxonomy, taxonomic classification system of angiosperms, flowering plants. It was developed by Arthur Cronquist in a series of monographs and texts, including ''The Evolution and Classification of ...
, but with somewhat greater complexity at the higher levels. He favors smaller orders and families, to allow character and evolutionary relationships to be more easily grasped. The Takhtajan classification system remains influential; it is used, for example, by the Montréal Botanical Garden.


Selected publications

* 1948. Морфологическая эволюция покрытосеменных (Morphological evolution of angiosperms). ranslated from Russian to German by Werner Höppner, as ''Die Evolution der Angiospermae'' 1959, Fischer, Jena * * 1969. Flowering plants: origin and dispersal. Oliver and Boyd, Edinburgh C Jeffrey">Charles_Jeffrey_(botanist).html" ;"title="ranslated from Russian by Charles Jeffrey (botanist)">C Jeffreyref name=T1966/> * 1987. Systema Magnoliophytorum. Leningrad. * 2009. Flowering Plants. Springer, New York


Legacy

Takhtajan has been considered one of the leading botanists of his time. He has been honoured in the naming of several plant genera. In 1980, botanist Vandika Ervandovna Avetisyan published ''Takhtajaniella'', which is a genus of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
from Transcausica, belonging to the family
Brassicaceae Brassicaceae () or (the older but equally valid) Cruciferae () is a medium-sized and economically important Family (biology), family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous pla ...
and it was named in his honour. Then in 1990, Nazarova published ''
Takhtajaniantha ''Takhtajaniantha'' is a genus of flowering plants in the tribe Cichorieae within the family Asteraceae, widespread across central Asia, Middle east and eastern Europe.Nazarova, Estella A. 1990. Biologicheskii Zhurnal Armenii 43: 179–183 The ...
'' (from the dandelion tribe within the daisy family,Nazarova, Estella A. 1990. Biologicheskii Zhurnal Armenii 43: 179–183) and lastly in 1997, '' Takhtajania'' (from the family
Winteraceae Winteraceae is a primitive family of tropical trees and shrubs including 93 species in five genera. It is of particular interest because it is such a primitive angiosperm family, distantly related to Magnoliaceae, though it has a much more south ...
, which was found in
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
) was published.World Conservation Monitoring Centre. 1998
''Takhtajania perrieri''.
The IUCN Red List of Threatened Species 1998. Downloaded on 10 October 2015.


See also

*
Takhtajan system A list of systems of plant taxonomy, system of plant taxonomy, the Takhtajan system of plant classification was published by Armen Takhtajan, in several versions from the 1950s onwards. It is usually compared to the Cronquist system. It admits Para ...


References


Works by Takhtajan

* * *Takhtajan, Armen. 1973. Chetyre tsarstva organicheskogo mira (Four Kingdoms of the Organic World). Priroda (Nature), Akad. Nauk 2: 22–32. * * * * * *


External links

*
The parting with Armen Takhtajan
Photo-report on the site of the Komarov Botanical Institute.

{{DEFAULTSORT:Takhtajan, Armen 1910 births 2009 deaths Russian people of Armenian descent Armenian botanists Full Members of the USSR Academy of Sciences Full Members of the Russian Academy of Sciences Phytogeographers Pteridologists Soviet botanists Scientists from Shusha Symbiogenesis researchers Foreign associates of the National Academy of Sciences Members of the German National Academy of Sciences Leopoldina Members of the Norwegian Academy of Science and Letters Yerevan State University alumni Academic staff of Yerevan State University Russian scientists Leipzig University alumni Soviet Armenians