Tachymetabolism
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Warm-blooded is a term referring to
animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
whose bodies maintain a temperature higher than that of their environment. In particular,
homeothermic Homeothermy, homothermy, or homoiothermy () is thermoregulation that maintains a stable internal body temperature regardless of external influence. This internal body temperature is often, though not necessarily, higher than the immediate envir ...
species (including
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s and
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s) maintain a stable body temperature by regulating
metabolic Metabolism (, from ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cellular processes; the ...
processes. Other species have various degrees of
thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
. Because there are more than two categories of temperature control utilized by animals, the terms ''warm-blooded'' and '' cold-blooded'' have been deprecated in the scientific field.


Terminology

In general, warm-bloodedness refers to three separate categories of
thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
. * Endothermy is the ability of some creatures to control their body temperatures through internal means such as muscle shivering or increasing their
metabolism Metabolism (, from ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cellular processes; the co ...
. The opposite of endothermy is
ectothermy An ectotherm (), more commonly referred to as a "cold-blooded animal", is an animal in which internal physiological sources of heat, such as blood, are of relatively small or of quite negligible importance in controlling body temperature.Daven ...
. * Homeothermy maintains a stable internal body temperature regardless of external influence and temperatures. The stable internal temperature is often higher than the immediate environment. The opposite is
poikilothermy A poikilotherm () is an animal (Greek ''poikilos'' – 'various', 'spotted', and ''therme'' – 'heat') whose internal temperature varies considerably. Poikilotherms have to survive and adapt to environmental stress. One of the most important s ...
. The only known living homeotherms are
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s and
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s, as well as one lizard, the
Argentine black and white tegu The Argentine black and white tegu (''Salvator merianae''), also Common name, commonly known as the Argentine giant tegu, the black and white tegu, the blue tegu, and the huge tegu, is a species of lizards in the Family (biology), family Teiidae ...
. Some non-avian
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s as well as some extinct reptiles such as
ichthyosaur Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides. Ichthyosaurians thrived during much of the Mesozoic era; based on fo ...
s,
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
s,
plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...
s are believed to have been homeotherms. *Tachymetabolism maintains a high "resting" metabolism. In essence, tachymetabolic creatures are "on" all the time. Though their resting metabolism is still many times slower than their active metabolism, the difference is often not as large as that seen in bradymetabolic creatures. Tachymetabolic creatures have greater difficulty dealing with a scarcity of food.


Varieties of thermoregulation

A significant proportion of creatures commonly referred to as "warm-blooded," like birds and mammals, exhibit all three of these categories (i.e., they are endothermic, homeothermic, ''and'' tachymetabolic). However, over the past three decades, investigations in the field of animal thermophysiology have unveiled numerous species within these two groups that do not meet all these criteria. For instance, many bats and small birds become poikilothermic and bradymetabolic during sleep (or, in nocturnal species, during the day). For such creatures, the term ''
heterothermy Heterothermy or heterothermia (from Greek ἕτερος ''heteros'' "other" and θέρμη ''thermē'' "heat") is a physiological term for animals that vary between self-regulating their body temperature, and allowing the surrounding environment t ...
'' was introduced. Further examinations of animals traditionally classified as cold-blooded have revealed that most creatures manifest varying combinations of the three aforementioned terms, along with their counterparts (ectothermy, poikilothermy, and bradymetabolism), thus creating a broad spectrum of body temperature types. Some
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
have warm-blooded characteristics, such as the
opah The opah, also commonly known as moonfish, sunfish (not to be confused with Molidae), cowfish, kingfish, and redfin ocean pan are large, colorful, deep-bodied Pelagic zone, pelagic lampriform Fish, fishes comprising the genus ''Lampris'', of ...
.
Swordfish The swordfish (''Xiphias gladius''), also known as the broadbill in some countries, are large, highly migratory predatory fish characterized by a long, flat, pointed bill. They are the sole member of the Family (biology), family Xiphiidae. They ...
and some
shark Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the ...
s have
circulatory In vertebrates, the circulatory system is a system of organs that includes the heart, blood vessels, and blood which is circulated throughout the body. It includes the cardiovascular system, or vascular system, that consists of the heart an ...
mechanisms that keep their
brain The brain is an organ (biology), organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It consists of nervous tissue and is typically located in the head (cephalization), usually near organs for ...
s and
eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...
s above ambient temperatures and thus increase their ability to detect and react to
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
. Tunas and some sharks have similar mechanisms in their muscles, improving their stamina when swimming at high speed.


Heat generation

Body heat is generated by
metabolism Metabolism (, from ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cellular processes; the co ...
. This relates to the chemical reaction in
cell Cell most often refers to: * Cell (biology), the functional basic unit of life * Cellphone, a phone connected to a cellular network * Clandestine cell, a penetration-resistant form of a secret or outlawed organization * Electrochemical cell, a de ...
s that break down
glucose Glucose is a sugar with the Chemical formula#Molecular formula, molecular formula , which is often abbreviated as Glc. It is overall the most abundant monosaccharide, a subcategory of carbohydrates. It is mainly made by plants and most algae d ...
into water and
carbon dioxide Carbon dioxide is a chemical compound with the chemical formula . It is made up of molecules that each have one carbon atom covalent bond, covalently double bonded to two oxygen atoms. It is found in a gas state at room temperature and at norma ...
, thereby producing
adenosine triphosphate Adenosine triphosphate (ATP) is a nucleoside triphosphate that provides energy to drive and support many processes in living cell (biology), cells, such as muscle contraction, nerve impulse propagation, and chemical synthesis. Found in all known ...
(ATP), a high-energy compound used to power other cellular processes. Muscle contraction is one such metabolic process generating heat energy, and additional heat results from friction as blood circulates through the vascular system in premise to their specialized fat
cells Cell most often refers to: * Cell (biology), the functional basic unit of life * Cellphone, a phone connected to a cellular network * Clandestine cell, a penetration-resistant form of a secret or outlawed organization * Electrochemical cell, a d ...
which produce heat through uncoupled
respiration Respiration may refer to: Biology * Cellular respiration, the process in which nutrients are converted into useful energy in a cell ** Anaerobic respiration, cellular respiration without oxygen ** Maintenance respiration, the amount of cellul ...
, contributing to
thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
. All organisms metabolize food and other inputs, but some make better use of the output than others. Like all energy conversions, metabolism is rather inefficient, and around 60% of the available energy is converted to heat rather than to ATP. In most organisms, this heat dissipates into the surroundings. However, endothermic homeotherms (generally referred to as "warm-blooded" animals) not only produce more heat but also possess superior means of retaining and regulating it compared to other animals. They exhibit a higher basal metabolic rate and can further increase their metabolic rate during strenuous activity. They usually have well-developed insulation in order to retain body heat: fur and
blubber Blubber is a thick layer of Blood vessel, vascularized adipose tissue under the skin of all cetaceans, pinnipeds, penguins, and sirenians. It was present in many marine reptiles, such as Ichthyosauria, ichthyosaurs and plesiosaurs. Description ...
in the case of mammals and
feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and an exa ...
s in birds. When this insulation is insufficient to maintain body temperature, they may resort to
shivering Shivering (also called shuddering) is a bodily function in response to cold and extreme fear in warm-blooded animals. When the core body temperature drops, the shivering reflex is triggered to maintain homeostasis. Skeletal muscles begin to sha ...
—rapid muscle contractions that quickly use up ATP, thus stimulating cellular metabolism to replace it and consequently produce more heat. Additionally, almost all
eutheria Eutheria (from Greek , 'good, right' and , 'beast'; ), also called Pan-Placentalia, is the clade consisting of Placentalia, placental mammals and all therian mammals that are more closely related to placentals than to marsupials. Eutherians ...
n mammals (with the only known exception being
swine Suina (also known as Suiformes) is a suborder of omnivorous, non-ruminant artiodactyl mammals that includes the domestic pig and peccaries. A member of this clade is known as a suine. Suina includes the family Suidae, termed suids, known in ...
) have
brown adipose tissue Brown adipose tissue (BAT) or brown fat makes up the adipose organ together with white adipose tissue (or white fat). Brown adipose tissue is found in almost all mammals. Classification of brown fat refers to two distinct cell populations with si ...
whose
mitochondria A mitochondrion () is an organelle found in the cells of most eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is us ...
are capable of non-shivering thermogenesis. This process involves the direct dissipation of the mitochondrial gradient as heat via an
uncoupling protein An uncoupling protein (UCP) is a mitochondrial inner membrane protein that is a regulated proton channel or transporter. An uncoupling protein is thus capable of dissipating the proton gradient generated by NADH-powered pumping of protons from the ...
, thereby "uncoupling" the gradient from its usual function of driving ATP production via
ATP synthase ATP synthase is an enzyme that catalyzes the formation of the energy storage molecule adenosine triphosphate (ATP) using adenosine diphosphate (ADP) and inorganic phosphate (Pi). ATP synthase is a molecular machine. The overall reaction catalyzed ...
. In warm environments, these animals employ evaporative cooling to shed excess heat, either through
sweating Perspiration, also known as sweat, is the fluid secreted by sweat glands in the skin of mammals. Two types of sweat glands can be found in humans: eccrine glands and apocrine glands. The eccrine sweat glands are distributed over much of the ...
(some mammals) or by panting (many mammals and all birds)—mechanisms generally absent in poikilotherms.


Defense against fungi

It has been hypothesized that warm-bloodedness evolved in mammals and birds as a defense against
fungal infection Fungal infection, also known as mycosis, is a disease caused by fungi. Different types are traditionally divided according to the part of the body affected: superficial, subcutaneous, and systemic. Superficial fungal infections include common ...
s. Very few fungi can survive the body temperatures of warm-blooded animals. By comparison, insects, reptiles, and amphibians are plagued by fungal infections. Warm-blooded animals have a defense against pathogens contracted from the environment, since environmental pathogens are not adapted to their higher internal temperature.


See also

* *
Mesotherm A mesotherm () is a type of animal with a thermoregulatory strategy intermediate to cold-blooded ectotherms and warm-blooded endotherms. Definition Mesotherms have two basic characteristics: # Elevation of body temperature via metabolic produ ...
*
Thermogenic plant Thermogenic plants have the ability to raise their temperature above that of the surrounding air. Heat is generated in the mitochondria, as a secondary process of cellular respiration called thermogenesis. Alternative oxidase and uncoupling protein ...


References

Footnotes Citations


External links


What is Warm Blooded??
{{DEFAULTSORT:Warm-Blooded Animal physiology Thermoregulation ca:Sang calenta it:Omeotermia pt:Homeotermia