Syringa Protolaciniata
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''Syringa'' is a genus of 12 currently recognized species of
flowering Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
woody plant A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to Herbaceous plant, herbaceous plants that die back to t ...
s in the olive family or
Oleaceae Oleaceae, also known as the olive family or sometimes the lilac family, is a taxonomic family of flowering shrubs, trees, and a few lianas in the order Lamiales. It presently comprises 28 genera, one of which is recently extinct.Peter S. Gree ...
called lilacs. These lilacs are native to woodland and scrub from southeastern
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
to eastern
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, and widely and commonly cultivated in temperate areas elsewhere.Flora Europaea
''Syringa''
/ref>Flora of China
丁香属 ding xiang shu ''Syringa''
/ref>Flora of Pakistan
''Syringa''
/ref>Germplasm Resources Information Network
''Syringa''
The genus is most closely related to ''
Ligustrum A privet is a flowering plant in the genus ''Ligustrum''. The genus contains about 50 species of erect, deciduous or evergreen shrubs or trees, with a native distribution from Europe to tropical and subtropical Asia, and with one species each ...
'' (privet), classified with it in Oleaceae tribus Oleeae subtribus Ligustrinae.University of Oxford, Oleaceae information site
New classification of the Oleaceae
/ref> Lilacs are used as food plants by the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of some moth species, including lilac leaf mining moth,
privet hawk moth ''Sphinx ligustri'', the privet hawk moth, is a moth found in most of the Palearctic realm. The species was Species description, first described by Carl Linnaeus in his 1758 10th edition of Systema Naturae, 10th edition of ''Systema Naturae''. ...
,
copper underwing The copper underwing, humped green fruitworm or pyramidal green fruitworm (''Amphipyra pyramidea'') is a moth of the family Noctuidae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. Distributi ...
,
scalloped oak The scalloped oak (''Crocallis elinguaria'') is a moth of the family Geometridae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. Distribution This common species can be found from Europe to eas ...
and
Svensson's copper underwing Svensson's copper underwing (''Amphipyra berbera'') is a moth of the family Noctuidae. The species was first described by Charles E. Rungs in 1949. It is distributed throughout Europe including Russia east to the Urals. This species has a wings ...
.


Description

Lilacs are small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s, ranging in size from tall, with stems up to diameter. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are opposite (occasionally in whorls of three) in
arrangement In music, an arrangement is a musical adaptation of an existing composition. Differences from the original composition may include reharmonization, melodic paraphrasing, orchestration, or formal development. Arranging differs from orchestr ...
, and their
shape A shape is a graphics, graphical representation of an object's form or its external boundary, outline, or external Surface (mathematics), surface. It is distinct from other object properties, such as color, Surface texture, texture, or material ...
is simple and heart-shaped to broad lanceolate in most species, but
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...
in a few species (e.g. ''S. protolaciniata'', ''S. pinnatifolia'').


Flowers

The
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s are produced in spring, each flower being in diameter with a four-lobed corolla, the corolla tube narrow, long; they are
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...
, with fertile
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s and stigma in each flower. The usual flower colour is a shade of purple (often a light purple or "
lilac ''Syringa'' is a genus of 12 currently recognized species of flowering woody plants in the olive family or Oleaceae called lilacs. These lilacs are native to woodland and scrub from southeastern Europe to eastern Asia, and widely and commonly ...
"), but white, pale yellow and pink, and even a dark burgundy color are also found. The flowers grow in large
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
s, and in several species have a strong fragrance. Flowering varies between mid spring to early summer, depending on the species. One particular cultivar, trademark ''Bloomerang'', first blooms in spring and then again late summer through fall.


Fruit

The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
is a dry, brown capsule, splitting in two at maturity to release the two winged
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s.


Etymology

The English common name "lilac" is from the French ''lilac'' via the from meaning the
indigo plant ''Indigofera'' is a large genus of over 750 species of flowering plants belonging to the pea family Fabaceae. They are widely distributed throughout the tropical and subtropical regions of the world. Description ''Indigofera'' is a varied genu ...
or ''nilak'' meaning "bluish"; both ''lilanj'' and ''nilak'' come from Persian ''nīl'' "
indigo InterGlobe Aviation Limited (d/b/a IndiGo), is an India, Indian airline headquartered in Gurgaon, Haryana, India. It is the largest List of airlines of India, airline in India by passengers carried and fleet size, with a 64.1% domestic market ...
" or ''nili'' "dark blue".


Taxonomy

The genus ''Syringa'' was first formally described in 1753 by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
and the description was published in ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genus, genera. It is the first work to consistently apply binomial nomenclature ...
''. The genus name ''Syringa'' is derived from
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
word ''syrinx'' meaning "pipe" or "tube" and refers to the hollow branches of '' S. vulgaris''. Homonym ''Syringa'' Tourn. ex Adans. is a heterotypic
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
of
Philadelphus ''Philadelphus'' () (mock-orange) is a genus of about 60 species of shrubs from 3–20 ft (1–6 m) tall, native to North America, Central America, Asia and (locally) in southeast Europe. They are named "mock-orange" in reference to their ...
.


Cultivation and uses

Lilacs are popular shrubs in
park A park is an area of natural, semi-natural or planted space set aside for human enjoyment and recreation or for the protection of wildlife or natural habitats. Urban parks are urban green space, green spaces set aside for recreation inside t ...
s and
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...
s throughout the temperate zone, and several
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...
s and numerous
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s have been developed. The term French lilac is often used to refer to modern
double-flowered "Double-flowered" describes varieties of flowers with extra petals, often containing flowers within flowers. The double-flowered trait is often noted alongside the scientific name with the abbreviation ''fl. pl.'' (''flore pleno'', a Latin ablati ...
cultivars, thanks to the work of prolific breeder
Victor Lemoine Pierre Louis Victor Lemoine (October 21, 1823 in Delme, Moselle - December 11, 1911) was a celebrated and prolific French flower breeder who, among other accomplishments, created many of today's lilac varieties. As a result of his accomplishmen ...
. Lilacs grow most successfully in well-drained soils, particularly those based on chalk. They flower on old wood, and produce more flowers if unpruned. If pruned, the plant responds by producing fast-growing young vegetative growth with no flowers, in an attempt to restore the removed branches. Lilac bushes can be prone to powdery mildew disease. Lilac wood is not commonly used or commercially harvested due to the small size of the tree. It is a relatively hard wood, with an estimated Janka hardness of 2,350 lbf (10,440 N), and is reportedly good for
woodturning Woodturning is the craft of using a wood lathe with hand-held tools to cut a shape that is symmetrical around the axis of rotation. Like the potter's wheel, the wood lathe is a mechanism that can generate a variety of forms. The operator is kno ...
. The sapwood is typically cream-coloured and the heartwood can have various streaks of brown and purple. Species have been historically used in various traditional medicines in Asia for treating ailments including cough, diarrhea, acute
icteric Jaundice, also known as icterus, is a yellowish or, less frequently, greenish pigmentation of the skin and sclera due to high bilirubin levels. Jaundice in adults is typically a sign indicating the presence of underlying diseases involving ...
hepatitis Hepatitis is inflammation of the liver parenchyma, liver tissue. Some people or animals with hepatitis have no symptoms, whereas others develop yellow discoloration of the skin and whites of the eyes (jaundice), Anorexia (symptom), poor appetite ...
, vomiting, abdominal pain, and bronchitis. Compounds isolated from species of ''Syringa'' include
phenylpropanoid The phenylpropanoids are a diverse family of organic compounds that are biosynthesized by plants from the amino acids phenylalanine and tyrosine in the shikimic acid pathway. Their name is derived from the six-carbon, aromatic phenyl group and ...
s such as syringin and
iridoid Iridoids are a type of monoterpenoids in the general form of cyclopentanopyran, found in a wide variety of plants and some animals. They are biosynthetically derived from 8-oxogeranial. Iridoids are typically found in plants as glycosides, mo ...
s such as
oleuropein Oleuropein is a glycosylation, glycosylated seco-iridoid, a type of polyphenol, phenolic bitter compound found in green olive skin, flesh, seeds, and leaves. The term oleuropein is derived from the botanical name of the olive tree, ''Olea europaea ...
. Substituent compounds, such as iridoids, as well as crude extracts from ''Syringa'' plants have been shown to have to have effects including
antitumor Cancer treatments are a wide range of treatments available for the many different types of cancer, with each cancer type needing its own specific treatment. Treatments can include surgery, chemotherapy, radiation therapy, hormonal therapy, targe ...
,
antihypertensive Antihypertensives are a class of drugs that are used to treat hypertension (high blood pressure). Antihypertensive therapy seeks to prevent the complications of high blood pressure, such as stroke, heart failure, kidney failure and myocardial infa ...
, anti-inflammatory, antioxidant, and antifungal activities in pharmacological studies.


Symbolism

Lilacs are often considered to symbolize first love. In
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
,
Macedonia Macedonia (, , , ), most commonly refers to: * North Macedonia, a country in southeastern Europe, known until 2019 as the Republic of Macedonia * Macedonia (ancient kingdom), a kingdom in Greek antiquity * Macedonia (Greece), a former administr ...
,
Lebanon Lebanon, officially the Republic of Lebanon, is a country in the Levant region of West Asia. Situated at the crossroads of the Mediterranean Basin and the Arabian Peninsula, it is bordered by Syria to the north and east, Israel to the south ...
, and
Cyprus Cyprus (), officially the Republic of Cyprus, is an island country in the eastern Mediterranean Sea. Situated in West Asia, its cultural identity and geopolitical orientation are overwhelmingly Southeast European. Cyprus is the List of isl ...
, the lilac is strongly associated with
Easter Easter, also called Pascha ( Aramaic: פַּסְחָא , ''paskha''; Greek: πάσχα, ''páskha'') or Resurrection Sunday, is a Christian festival and cultural holiday commemorating the resurrection of Jesus from the dead, described in t ...
time because it flowers around that time; it is consequently called ''paschalia''. In the poem ''
When Lilacs Last in the Dooryard Bloom'd "When Lilacs Last in the Dooryard Bloom'd" is a long poem written by American poet Walt Whitman (1819–1892) as an elegy to President Abraham Lincoln. It was written in the summer of 1865 during a period of profound national mourning in the ...
'', by
Walt Whitman Walter Whitman Jr. (; May 31, 1819 – March 26, 1892) was an American poet, essayist, and journalist; he also wrote two novels. He is considered one of the most influential poets in American literature and world literature. Whitman incor ...
, lilacs are a reference to
Abraham Lincoln Abraham Lincoln (February 12, 1809 – April 15, 1865) was the 16th president of the United States, serving from 1861 until Assassination of Abraham Lincoln, his assassination in 1865. He led the United States through the American Civil War ...
. The music-hall song by
Ivor Novello Ivor Novello (born David Ivor Davies; 15 January 1893 – 6 March 1951) was a Welsh actor, dramatist, singer and composer who became one of the most popular British entertainers of the first half of the 20th century. He was born into a musical ...
, '' We'll Gather Lilacs'', first performed in 1945, speaks of the longing of two lovers to be reunited in a traditional English rural setting. It has since been recorded and performed by numerous artists. ''Syringa vulgaris'' is the
state flower This is a list of U.S. state, federal district, and territory flowers. See also *List of U.S. state trees * Lists of U.S. state insignia References External linksList of state flowers {{USStateLists * U.S. state flowers Flowers ...
of
New Hampshire New Hampshire ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Massachusetts to the south, Vermont to the west, Maine and the Gulf of Maine to the east, and the Canadian province of Quebec t ...
, because it "is symbolic of that hardy character of the men and women of the Granite State."


Festivals

Several locations in North America hold annual Lilac Festivals, including: * The
Arnold Arboretum The Arnold Arboretum is a botanical research institution and free public park affiliated with Harvard University and located in the Jamaica Plain and Roslindale, Massachusetts, Roslindale neighborhoods of Boston. Established in 1872, it is the ...
in
Boston Boston is the capital and most populous city in the Commonwealth (U.S. state), Commonwealth of Massachusetts in the United States. The city serves as the cultural and Financial centre, financial center of New England, a region of the Northeas ...
,
Massachusetts Massachusetts ( ; ), officially the Commonwealth of Massachusetts, is a U.S. state, state in the New England region of the Northeastern United States. It borders the Atlantic Ocean and the Gulf of Maine to its east, Connecticut and Rhode ...
, which celebrates "Lilac Sunday" every May. The Arboretum shows off its collection of over 422 lilac plants, of 194 different varieties. Lilac Sunday is the only day of the year when picnicking is allowed on the grounds of the Arboretum. *
Lombard, Illinois Lombard is a village in DuPage County, Illinois, United States, and a suburb of Chicago. The population was 44,476 at the 2020 United States census, 2020 census. History Lombard was originally named "Babcock's Grove", after the Babcock brothers ...
, called the "Lilac Village", which has an annual lilac festival and parade in May. The village also contains Lilacia Park, a garden with over 200 varieties of lilacs, as well as over 50 kinds of
tulip Tulips are spring-blooming perennial herbaceous bulbiferous geophytes in the ''Tulipa'' genus. Their flowers are usually large, showy, and brightly coloured, generally red, orange, pink, yellow, or white. They often have a different colour ...
s. *
Mackinac Island Mackinac Island ( , ; ; ; ) is an island and resort area, covering in land area, in the U.S. state of Michigan. The name of the island in Odawa is Michilimackinac and "Mitchimakinak" in Ojibwemowin, meaning "Great Turtle". It is located in ...
, in
Michigan Michigan ( ) is a peninsular U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest, Upper Midwestern United States. It shares water and land boundaries with Minnesota to the northwest, Wisconsin to the west, ...
, which celebrates a weeklong lilac festival and lilac parade each June. *
Rochester, New York Rochester is a city in and the county seat, seat of government of Monroe County, New York, United States. It is the List of municipalities in New York, fourth-most populous city and 10th most-populated municipality in New York, with a populati ...
, which has held its Lilac Festival since 1898, hosts the longest-running festival in North America. Held in Highland Park, this celebration features 1,200 shrubs, representing over 500 varieties, many of which were developed in Rochester. It is the largest collection of varieties at any single place. * The Royal Botanical Gardens near
Hamilton Hamilton may refer to: * Alexander Hamilton (1755/1757–1804), first U.S. Secretary of the Treasury and one of the Founding Fathers of the United States * ''Hamilton'' (musical), a 2015 Broadway musical by Lin-Manuel Miranda ** ''Hamilton'' (al ...
,
Ontario Ontario is the southernmost Provinces and territories of Canada, province of Canada. Located in Central Canada, Ontario is the Population of Canada by province and territory, country's most populous province. As of the 2021 Canadian census, it ...
, which holds its Lilac Celebration each May. *
Spokane, Washington Spokane ( ) is the most populous city in eastern Washington and the county seat of Spokane County, Washington, United States. It lies along the Spokane River, adjacent to the Selkirk Mountains, and west of the Rocky Mountain foothills, south o ...
, known as the "Lilac City", which holds an annual lilac festival and lilac parade. * Franktown, Ontario, Canada, known as the Lilac Capital of Canada, holds an annual festival. With drystone masonry demonstrations and horse pulled wagon rides. *
Calgary Calgary () is a major city in the Canadian province of Alberta. As of 2021, the city proper had a population of 1,306,784 and a metropolitan population of 1,481,806 making it the third-largest city and fifth-largest metropolitan area in C ...
, Alberta, Canada, holds an annual one-day Lilac Festival, which is primarily a street festival.


Species

Species and subspecies currently accepted as of July 2016: *'' Syringa emodi'' Wall. ex Royle – Himalayan lilac - northern India, Pakistan, Tibet, Nepal *'' Syringa josikaea'' J.Jacq. ex Rchb.f. – Hungarian lilac - Carpathian Mountains of Romania and Ukraine *'' Syringa komarowii'' C.K.Schneid. – nodding lilac - Gansu, Hubei, Shaanxi, Sichuan, Yunnan *'' Syringa oblata'' Lindl. – early blooming lilac or broadleaf lilac - Korea, Gansu, Hebei, Henan, Jilin, Liaoning, Inner Mongolia, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan **'' ''Syringa oblata'' subsp. ''dilatata'' – Korean early lilac - Nakai - Korea, Jilin, Liaoning *'' Syringa pinetorum'' W.W.Sm. – Sichuan, Tibet, Yunnan *'' Syringa pinnatifolia'' Hemsl. – Gansu, Inner Mongolia, Ningxia, Qinghai, Shaanxi, Sichuan *'' Syringa pubescens'' Turcz. – Korea, Gansu, Hebei, Henan, Hubei, Jilin, Liaoning, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan *'' Syringa reticulata'' (Blume) H.Hara (syn. ''S. pekinensis'') – Japanese tree lilac - Primorye, Japan, Korea, Gansu, Hebei, Heilongjiang, Henan, Jilin, Liaoning, Inner Mongolia, Ningxia, Shaanxi, Shanxi, Sichuan *'' Syringa tomentella'' Bureau & Franch. – Sichuan, Tibet, Yunnan *'' Syringa villosa'' Vahl – villous lilac - Primorye, Korea, Hebei, Shanxi, Heilongjiang, Jilin, Liaoning *''
Syringa vulgaris ''Syringa vulgaris'', the lilac or common lilac, is a species of flowering plant in the olive family, Oleaceae. Native to the Balkan Peninsula, it is widely cultivated for its scented flowers in Europe (particularly the north and west) and Nor ...
'' L. – common lilac - native to Balkans; naturalized in western and central Europe, and many scattered locations in North America


Hybrids

* ''S.'' × ''chinensis'' (''S. vulgaris'' × ''S. persica'') * ''S.'' × ''diversifolia'' (''S. oblata'' × ''S. pinnatifolia'') *''S.'' × ''henryi'' (''S. josikaea'' × ''S. villosa'') *''S.'' × ''hyacinthiflora'' (''S. oblata'' × ''S. vulgaris'') *''S.'' × ''josiflexa'' (''S. josikaea'' × ''S. komarowii'') * ''S.'' × ''laciniata'' (''S. protolaciniata'' × ''S. vulgaris'') – cut-leaf lilac or cutleaf lilac * ''S.'' × ''persica'' L. (syn ''Syringa protolaciniata'') – Persian lilac - Afghanistan, Pakistan, western Himalayas, Gansu, Qinghai *''S.'' × ''prestoniae'' (''S. komarowii'' × ''S. villosa'') *''S.'' × ''swegiflexa'' (''S. komarowii'' × ''S. sweginzowii'')


Gallery

File:Syringa microphylla C.jpg, ''Syringa microphylla'' File:Syringa emodi 002.jpg, ''Syringa emodi'' flowers File:Syringa vulgaris Sarah Sands.jpg, ''Syringa vulgaris'' 'Sarah Sands' File:Syringa meyeri Palibin.jpg, ''Syringa meyeri'' 'Palibin' File:Syringa 'Pamyat o Vekhove' 01.jpg, Syringa 'Pamyat o Vekhove' File:Ab plant 487.jpg, White syringa File:Šeřík1.jpg, Syringarium with trees in blossom File:Syringa 'Pavlinka' 03.JPG, Syringa 'Pavlinka' File:Syringa 'Oberon' 01.JPG, Syringa 'Oberon' File:Lilacs 2.jpg, A double-flowered ''Syringa vulgaris'' cultivar


References


External links

{{Authority control Garden plants Oleaceae genera Shrubs Taxa named by Carl Linnaeus