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''Symphyotrichum dumosum'' (formerly ''Aster dumosus'') is a species of flowering plant of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...
commonly known as rice button aster and bushy aster. It is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to much of eastern and central North America, as well as
Haiti Haiti (; ht, Ayiti ; French: ), officially the Republic of Haiti (); ) and formerly known as Hayti, is a country located on the island of Hispaniola in the Greater Antilles archipelago of the Caribbean Sea, east of Cuba and Jamaica, and ...
and
Dominican Republic The Dominican Republic ( ; es, República Dominicana, ) is a country located on the island of Hispaniola in the Greater Antilles archipelago of the Caribbean region. It occupies the eastern five-eighths of the island, which it shares wit ...
. It is a
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
,
herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
that may reach a height of .


Description

''Symphyotrichum dumosum'' is a
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
,
herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
that grows up to high.


Flowers

''Symphyotrichum dumosum'' is a late-summer and fall blooming perennial, with flower heads opening August–October. The
flower heads A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
are about diameter when in bloom and grow in open and much-branched paniculiform arrays. Each has many tiny
florets This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
put together into what appear as one.


Involucres and phyllaries

On the outside the flower heads of all members of the family Asteraceae are small bracts that look like scales. These are called '' phyllaries'', and together they form the involucre that protects the individual flowers in the head before they open. The involucres of ''Symphyotrichum dumosum'' are cylinder-bell in shape and usually long. The phyllaries are
appressed This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
or slightly spreading. The shape of the outer phyllaries is oblong-oblanceolate or linear-oblanceolate, and the innermost phyllaries are linear. They are in strongly unequal rows, meaning they are very staggered and do not end at the same point, and they are hairless.


Florets

The 15–33 ray florets are blue, pink, purple, or white, with a length averaging and width of . Ray florets in the ''Symphyotrichum'' genus are exclusively female, each having a
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
(with style, stigma, and
ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
) but no
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
; thus, ray florets accept
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
and each can develop a
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
, but they produce no pollen. The 15–30 disk florets start out as pale yellow and turn pink with age. They are in the shape of a narrow funnel and are shallow at in depth. The disk florets are each made up of , collectively a '' corolla'', which open into 5 lanceolate lobes comprising less than of the depth of the floret. Disk florets in the ''Symphyotrichum'' genus are
bisexual Bisexuality is a romantic or sexual attraction or behavior toward both males and females, or to more than one gender. It may also be defined to include romantic or sexual attraction to people regardless of their sex or gender identity, whi ...
, each with both male (stamen,
anther The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
s, and filaments) and female reproductive parts; thus, a disk floret produces pollen and can develop a seed. Rice button aster (Symphyotrichum dumosum) (6435495559).jpg, Flower head showing ray and disk florets Symphyotrichum dumosum 107676031.jpg, Involucre, phyllaries, and bracts Symphyotrichum dumosum 108257760.jpg, Involucre, phyllaries, and bracts


Chromosomes

''Symphyotrichum dumosum'' has a chromosome number of ''x'' = 8.
Diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
and tetraploid
cytotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
s with respective chromosome counts of 16 and 32 have been reported.


Taxonomy

This species was long known as ''Aster dumosus'' before it was moved into its current genus due to a
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
analysis performed by American botanist
Guy L. Nesom Guy L. Nesom (born August 2, 1945)''U.S. Public Records Index'' Vol 2 (Provo, UT: Ancestry.com Operations, Inc.), 2010. is an American writer and botanist. Nesom received his Ph.D. in systematic botany from the University of North Carolina in 19 ...
. The genus ''Aster'' has been narrowed considerably in scope in recent years, due to new information about the relationships of the species in the group. ''Symphyotrichum dumosum'' is classified in the
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
''Symphyotrichum'', section ''Symphyotrichum'', subsection ''Dumosi''. It is one of the "bushy asters and relatives." The word ''Symphyotrichum'' has as its root the Greek , which means "coming together," and , which means "hair." The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
(second part of the
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
) is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
neuter case of and means "full of thornbushes" and "bushy" (rare).


Infraspecies

Although the following infraspecies are no longer accepted varieties according to
Catalogue of Life The Catalogue of Life is an online database that provides an index of known species of animals, plants, fungi, and microorganisms. It was created in 2001 as a partnership between the global Species 2000 and the American Integrated Taxonomic Info ...
(COL),
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...
(POWO), and World Flora Online (WFO), they were accepted by USDA PLANTS Database, NatureServe, and
Integrated Taxonomic Information System The Integrated Taxonomic Information System (ITIS) is an American partnership of federal agencies designed to provide consistent and reliable information on the taxonomy of biological species. ITIS was originally formed in 1996 as an interagenc ...
(ITIS). The autonym is ''Symphyotrichum dumosum'' var. ''dumosum''. *''Symphyotrichum dumosum'' var. ''gracilipes'' *''Symphyotrichum dumosum'' var. ''pergracile'' *''Symphyotrichum dumosum'' var. ''strictior'' *''Symphyotrichum dumosum'' var. ''subulifolium''


Distribution and habitat


Distribution

''Symphyotrichum dumosum'' has a recorded native presence in the wild in the
Canadian Canadians (french: Canadiens) are people identified with the country of Canada. This connection may be residential, legal, historical or cultural. For most Canadians, many (or all) of these connections exist and are collectively the source of ...
province of
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
, and in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
in all states east of the
Mississippi River The Mississippi River is the second-longest river and chief river of the second-largest drainage system in North America, second only to the Hudson Bay drainage system. From its traditional source of Lake Itasca in northern Minnesota, it f ...
except
Vermont Vermont () is a state in the northeast New England region of the United States. Vermont is bordered by the states of Massachusetts to the south, New Hampshire to the east, and New York to the west, and the Canadian province of Quebec to ...
, as well as west of the Mississippi River in
Iowa Iowa () is a state in the Midwestern region of the United States, bordered by the Mississippi River to the east and the Missouri River and Big Sioux River to the west. It is bordered by six states: Wisconsin to the northeast, Illinois to the ...
,
Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ...
,
Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...
,
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...
,
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
, and
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
. It is also native on the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
island of
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. It is an
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in
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, and in the countries of
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,
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,
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,
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,
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
, and
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
. It is not on the European Union's
List of invasive alien species of Union concern In 2016, following the EU Regulation 1143/2014 on Invasive Alien Species (IAS), the European Commission published a first list of 37 IAS of Union concern. The list was first updated in 2017 and comprised 49 species. Since the second update in 201 ...
.


Habitat

''Symphyotrichum dumosum'' is categorized on the United States National Wetland Plant List (NWPL) with Wetland Indicator Status Rating of
Facultative {{wiktionary, facultative Facultative means "optional" or "discretionary" (antonym '' obligate''), used mainly in biology in phrases such as: * Facultative (FAC), facultative wetland (FACW), or facultative upland (FACU): wetland indicator statuses ...
(FAC), choosing wetlands or non-wetlands and adjusting accordingly.


Conservation

, NatureServe lists ''Symphyotrichum dumosum'' as Secure (G5) worldwide with notation that the global status was last reviewed in 1985. Some North American province and state statuses are as follows: Presumed Extirpated (SX) in
Iowa Iowa () is a state in the Midwestern region of the United States, bordered by the Mississippi River to the east and the Missouri River and Big Sioux River to the west. It is bordered by six states: Wisconsin to the northeast, Illinois to the ...
, Possibly Extirpated (SH) in
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and north ...
, Critically Imperiled (S1) in
Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...
, Imperiled (S2) in
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
and
Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...
, and Vulnerable (S3) in
New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...
and
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
.


Gallery

Mason Wasp on Bushy Aster (5250200700).jpg, Mason wasp on flower head Symphyotrichum dumosum 107676048.jpg, Stem and flower head Honey Bee on Symphyotrichum dumosum (6435501395).jpg, Honey bee on flower head Symphyotrichum dumosum 105499607.jpg, Cluster or small colony of ''S. dumosum''


Notes


Citations


References

* * * * * * * * * * * * * * * * * * {{Taxonbar, from1=Q4050490, from2=Q7061674 dumosum Flora of the United States Flora of Ontario Plants used in traditional Native American medicine Plants described in 1753 Taxa named by Carl Linnaeus