''Symphurus'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

mainly occur in relatively shallow water, including

estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...

. Some species are also found in

deeper water ''Deeper Water'', the tenth studio album by Paul Kelly, was released on 12 September 1995 on White Label Records in Australia and on Vanguard Records in North America. It peaked at No. 40 on the ARIA Albums Chart and provided his second c ...

, including '' S. thermophilus'' that lives at

hydrothermal vent Hydrothermal vents are fissures on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hot ...

s (the only

flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish Order (biology), suborder Pleuronectoidei, also called the Heterosomata. In many species, both eyes lie on one side of the head, one or the other migrating through or around ...

known from this habitat).Tyler, J. (2005): Distribution, population characteristics and trophic ecology of a sulphophilic hydrothermal vent tonguefish (Pleuronectiformes: Cynoglossidae). ''Master of Science. Thesis. University of Victoria: Canada.''Munroe, T.A. & Hashimoto, J. (2008): A new Western Pacific Tonguefish (Pleuronectiformes: Cynoglossidae): The first Pleuronectiform discovered at active Hydrothermal Vents. ''Zootaxa, 1839: 43–59.'' These species are distinguished by merged

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...

caudal Caudal may refer to: Anatomy * Caudal (anatomical term) (from Latin ''cauda''; tail), used to describe how close something is to the trailing end of an organism * Caudal artery, the portion of the dorsal aorta of a vertebrate that passes into th ...

and

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

s, the absence of a

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

and

pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...

s, and the presence of only one

pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...

. They are sinistral flatfishes, meaning that as

adult An adult is an animal that has reached full growth. The biological definition of the word means an animal reaching sexual maturity and thus capable of reproduction. In the human context, the term ''adult'' has meanings associated with social an ...

s, their crania are asymmetrical, with both

eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...

s on the left side. The largest species grows to about long.

Species

There are currently 78 recognized species in this genus: * ''

Symphurus arawak ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' C. R. Robins & J. E. Randall, 1965 (Caribbean tonguefish) * ''

Symphurus atramentatus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

D. S. Jordan David Starr Jordan (January 19, 1851 – September 19, 1931) was the founding president of Stanford University, serving from 1891 to 1913. He was an ichthyologist during his research career. Prior to serving as president of Stanford Universi ...

& Bollman, 1890 (Ink-spot tonguefish) * ''

Symphurus atricauda ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' (

& C. H. Gilbert, 1880) (California tonguefish) * ''

Symphurus australis ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' McCulloch, 1907 * ''

Symphurus bathyspilus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' Krabbenhoft &

Munroe Munroe is a derivation of the Scottish surname Munro (surname), Munro, and may refer to: In politics * Allen Munroe (1819–1884), New York politician * Daniel Munroe Forney, Congressional Representative from North Carolina * George H. Munroe (18 ...

, 2003 * '' Symphurus billykrietei''

, 1998 (Kriete's tonguefish) * '' Symphurus callopterus''

Mahadeva Mahadeva may refer to: Religion *Mahadeva, a title of the Hindu god Shiva **Parashiva, a form of Shiva ** Parameshwara (god), a form of Shiva * Para Brahman, a Hindu deity *Adi-Buddha, in Buddhism, the "First Buddha" or the "Primordial Buddha" * ...

, 1989 (Chocolate tonguefish) * '' Symphurus caribbeanus''

, 1991 * '' Symphurus chabanaudi''

, 1990 (Chabanaud's tonguefish) * '' Symphurus civitatium'' Ginsburg, 1951 (Offshore tonguefish) * ''

Symphurus diabolicus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

, 1990 (Devil's tonguefish) * ''

Symphurus diomedeanus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' ( Goode & T. H. Bean, 1885) (Spotted-fin tonguefish) * '' Symphurus elongatus'' ( Günther, 1868) (Elongated tonguefish) * ''

Symphurus fasciolaris ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' C. H. Gilbert, 1892 (Banded tonguefish) * ''

Symphurus fuscus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' A. B. Brauer, 1906 * ''

Symphurus gilesii ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' (

Alcock Alcock is a surname. Notable people with the surname include: * Alfred William Alcock, British naturalist * C. W. Alcock, British sports administrator, creator of the FA Cup * Charles R. Alcock, American astronomer * Deborah Alcock, British auth ...

, 1889) * ''

Symphurus ginsburgi ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' Menezes & Benvegnú, 1976 (Ginsburg's tonguefish) * ''

Symphurus gorgonae ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' Chabanaud, 1948 (Gorgonian tonguefish) * '' Symphurus holothuriae'' Chabanaud, 1948 * ''

Symphurus hondoensis ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' C. L. Hubbs, 1915 * ''

Symphurus insularis ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

, Brito & C. Hernández, 2000 * '' Symphurus jenynsi'' Evermann & Kendall, 1906 (Jenyn's tonguefish) * '' Symphurus kyaropterygium'' Menezes & Benvegnú, 1976 * '' Symphurus leei''

& Bollman, 1890 (Lee's tonguefish) * '' Symphurus leucochilus'' M. Y. Lee,

& K. T. Shao, 2014 Lee, M.-Y., Munroe, T.A. & Shao, K.-T. (2014): Description of a new cryptic, shallow-water tonguefish (Pleuronectiformes: Cynoglossidae: ''Symphurus'') from the western North Pacific Ocean. ''Journal of Fish Biology, 85 (3): 563–585.'' * '' Symphurus ligulatus'' ( Cocco, 1844) * ''

Symphurus longirostris ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' M. Y. Lee,

& Y. Kai, 2016 (Long-snout tonguefish) Lee, M.-Y., Munroe, T.A. & Kai, Y. (2016): Description of a new cryptic species of tonguefish (Pleuronectiformes: Cynoglossidae: ''Symphurus'') from shallow waters off Japan. ''Ichthyological Research, 64 (1): 71–83.'' * '' Symphurus lubbocki''

, 1990 * '' Symphurus luzonensis'' Chabanaud, 1955 * ''

Symphurus macrophthalmus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

Norman Norman or Normans may refer to: Ethnic and cultural identity * The Normans, a people partly descended from Norse Vikings who settled in the territory of Normandy in France in the 9th and 10th centuries ** People or things connected with the Norma ...

, 1939 * '' Symphurus maculopinnis''

, J. Tyler & Tunnicliffe, 2011 Munroe, T.A., Tyler, J. & Tunnicliffe, V. (2011): Description and biological observations on a new species of deepwater symphurine tonguefish (Pleuronectiformes: Cynoglossidae: ''Symphurus'') collected at Volcano–19, Tonga Arc, West Pacific Ocean. ''Zootaxa, 3061: 53-66.'' * ''

Symphurus maldivensis ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' Chabanaud, 1955 * '' Symphurus marginatus'' ( Goode & T. H. Bean, 1886) (Margined tonguefish) * ''

Symphurus marmoratus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' Fowler, 1934 * '' Symphurus megasomus'' M. Y. Lee, H. M. Chen & K. T. Shao, 2009 (Giant tonguefish) * '' Symphurus melanurus'' H. W. Clark, 1936 (Drab tonguefish) * ''

Symphurus melasmatotheca ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

& Nizinski, 1990 (Black-stripe tonguefish) * '' Symphurus microlepis''

Garman Garman is a surname or first name. Notable people with the name include: Sports * Ann Garman, All-American Girls Professional Baseball League player * Judi Garman (born 1954), American softball coach * Mike Garman (born 1949), American baseball pla ...

, 1899 (Small-fin tonguefish) * '' Symphurus microrhynchus'' ( M. C. W. Weber, 1913) * '' Symphurus minor'' Ginsburg, 1951 (Large-scale tonguefish) * ''

Symphurus monostigmus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

, 2006 * ''

Symphurus multimaculatus ''Symphurus'' is a genus of fish in the family (biology), family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in Abyssal zon ...

'' M. Y. Lee,

& H. M. Chen, 2009 (Pepper-dot tonguefish) * '' Symphurus nebulosus'' ( Goode & T. H. Bean, 1883) (Freckled tonguefish) * ''

Symphurus nigrescens ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

Rafinesque Constantine Samuel Rafinesque-Schmaltz (; 22 October 178318 September 1840) was a French early 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ult ...

, 1810 (Tonguefish) * ''

Symphurus normani ''Symphurus'' is a genus of fish in the family (biology), family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in Abyssal zon ...

'' Chabanaud, 1950 (Norman's tonguefish) * '' Symphurus novemfasciatus'' S. C. Shen & W. W. Lin, 1984 * '' Symphurus ocellaris''

& D. R. Robertson, 2005 (Ring-tail tonguefish) * ''

Symphurus ocellatus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' von Bonde, 1922 (Double-spot tonguefish) * ''

Symphurus oculellus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

, 1991 * '' Symphurus oligomerus''

, 1990 (Spot-fin tonguefish) * ''

Symphurus ommaspilus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' J. E. Böhlke, 1961 (Ocellated tonguefish) * ''

Symphurus orientalis ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' ...

'' ( Bleeker, 1879) * '' Symphurus parvus'' Ginsburg, 1951 (Pygmy tonguefish) * ''

Symphurus pelicanus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' Ginsburg, 1951 (Long-tail tonguefish) * ''

Symphurus piger ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' ( Goode & T. H. Bean, 1886) (Deep-water tonguefish) * '' Symphurus plagiusa'' (

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

, 1766) (Black-cheek tonguefish) * '' Symphurus plagusia'' (

Bloch Bloch is a surname of German origin. Notable people with this surname include: A *Adele Bloch-Bauer (1881–1925), Austrian entrepreneur *Albert Bloch (1882–1961), American painter *Alexandre Bloch (1857–1919), French painter *Alfred Bloch ( ...

& J. G. Schneider, 1801) (Dusky-cheek tonguefish) * '' Symphurus prolatinaris''

, Nizinski &

, 1991 (Half-striped tonguefish) * '' Symphurus pusillus'' ( Goode & T. H. Bean, 1885) (Northern tonguefish) * '' Symphurus regani'' M. C. W. Weber & de Beaufort, 1929 * '' Symphurus reticulatus''

, 1990 * '' Symphurus rhytisma'' J. E. Böhlke, 1961 (Patch-tail tonguefish) * '' Symphurus schultzi'' Chabanaud, 1955 * '' Symphurus septemstriatus'' (

, 1891) (Seven-band tonguefish) * '' Symphurus stigmosus''

, 1998 (Blotch-fin tonguefish) * ''

Symphurus strictus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' C. H. Gilbert, 1905 (Black-belly tonguefish) * '' Symphurus tessellatus'' ( Quoy &

Gaimard Joseph Paul Gaimard (31 January 1793 – 10 December 1858) was a French naval surgeon and naturalist. Biography Gaimard was born at Saint-Zacharie on January 31, 1793. He studied medicine at the naval medical school in Toulon, subsequent ...

, 1824) * ''

Symphurus thermophilus ''Symphurus thermophilus'' is a species of tonguefish notable for being the only flatfish known to be an obligate inhabitant of hydrothermal vents. It is known to inhabit several widely dispersed locations in the western Pacific Ocean and occurs ...

& Hashimoto, 2008 (Western Pacific tonguefish) * '' Symphurus trewavasae'' Chabanaud, 1948 (Trewavas' tonguefish) * ''

Symphurus trifasciatus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' (

, 1894) (Three-band tonguefish) * '' Symphurus undatus'' C. H. Gilbert, 1905 * '' Symphurus undecimplerus''

& Nizinski, 1990 (Dark-cheek tonguefish) * ''

Symphurus urospilus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' Ginsburg, 1951 (Spot-tail tonguefish) * '' Symphurus vanmelleae'' Chabanaud, 1952 (Vanmelle's tonguefish) * ''

Symphurus variegatus ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' ( Gilchrist, 1903) * ''

Symphurus varius ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

, 1899 (Mottled tonguefish) * '' Symphurus williamsi''

& Culver, 1895 (William's tonguefish) * ''

Symphurus woodmasoni ''Symphurus'' is a genus of fish in the family Cynoglossidae found in the Atlantic, Indian and Pacific Ocean. Most species mainly occur in relatively shallow water, including estuaries. Some species are also found in deeper water, including '' S ...

'' (

, 1889)

References

{{Taxonbar, from=Q2785579 Cynoglossidae Taxa named by Constantine Samuel Rafinesque Marine fish genera