Syma Megarhyncha
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The mountain kingfisher (''Syma megarhyncha'') is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the subfamily
Halcyoninae The tree kingfishers, also called wood kingfishers or Halcyoninae, are the most numerous of the three subfamilies of birds in the kingfisher family, with around 70 species divided into 12 genera, including several species of kookaburras. The sub ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Alcedinidae. Adult males are long, and have a
rufous Rufous () is a color that may be described as reddish-brown or brownish- red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a d ...
head and underparts, greenish-blue upperparts, a dark blue tail, and black
flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...
s. They also have dark neck patches and loral patches. Females have dark crowns and the neck patches join at the
nape The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , ). The corresponding adjective is ''nuchal'', as in the term ''nuchal rigidity'' ...
. It is similar to the yellow-billed kingfisher, but can be distinguished by its larger size and a proportionally larger bill, along with a dark ridge along its culmen. It is endemic to
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
, where it is found through most of the
New Guinea Highlands The New Guinea Highlands, also known as the Central Range or Central Cordillera, is a long chain of mountain ranges on the island of New Guinea, including the island's highest peak, Puncak Jaya, Indonesia, , the highest mountain in Oceania. The r ...
, except for the
Bird's Head Peninsula The Bird's Head Peninsula ( Indonesian: , , meaning Bird's Head in Indonesian and Dutch) or Doberai Peninsula (''Semenanjung Doberai'') is a large peninsula that makes up the northwest portion of the island of New Guinea, comprising the Indones ...
. Its natural
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s are
subtropical or tropical moist lowland forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in large ...
and subtropical or tropical moist
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...
. It is found at elevations between , although it has also been reported as low as . It feeds on small
lizards Lizard is the common name used for all squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The ...
,
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e, and
insects Insects (from Latin ') are hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed ...
. It lays eggs in December, and eggs are laid in
clutches A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does the ...
of two. The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
classifies it as a least-concern species.


Taxonomy and systematics

The mountain kingfisher was described as ''Syma megarhycha'' by
Tommaso Salvadori Count Adelardo Tommaso Salvadori Paleotti (30 September 1835 – 9 October 1923) was an Italian zoologist and ornithologist. Biography Salvadori was born in Porto San Giorgio, son of Count Luigi Salvadori and Ethelyn Welby, who was English. His ...
in 1896, on the basis of specimens collected in 1893 in Moroka,
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
. The generic name ''Syma'' is from the name of a sea nymph in
Greek mythology Greek mythology is the body of myths originally told by the Ancient Greece, ancient Greeks, and a genre of ancient Greek folklore, today absorbed alongside Roman mythology into the broader designation of classical mythology. These stories conc ...
. The specific name come from the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
''megas'' (great) and ''rhunkhos'' (bill). Mountain kingfisher has been designated as the official
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
by the
International Ornithologists' Union The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", ...
(IOC). The species is also known as the greater yellow-billed kingfisher or mountain yellow-billed kingfisher. It is one of two species in the genus ''
Syma ''Syma'' is a genus of tree kingfishers in the family Alcedinidae that are resident in New Guinea and northeast Australia. The genus was introduced by the French surgeon and naturalist René Lesson in 1827. ''Syma'' was the name of a sea nymph ...
'', along with the yellow-billed kingfisher. These two species are closely related and were previously considered
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
. Hybrids between them have also been recorded.


Subspecies

There are two recognised subspecies of the mountain kingfisher: * ''S. m. megarhyncha'' Salvadori, 1896: The
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
, it is found western, central, and northeastern New Guinea. Populations from western New Guinea, the
Snow Mountains The Sudirman Range, also known as the Snow Mountains,"Sudirman Mountains (Snow Mountains)". Papua Insects Foundation. Accessed 5 August 2021/ref> Dugunduguoo, or Nassau Range is a mountain range in Central Papua province, Indonesia. It is named a ...
, and the Weyland Mountains are sometimes separated as ''S. m. wellsi'', on the basis of slight differences in body and bill size. * ''S. m. sellamontis'' Reichenow, 1919: Found in northeastern New Guinea, in the mountains of the
Huon peninsula Huon Peninsula is a large rugged peninsula on the island of New Guinea in Morobe Province, eastern Papua New Guinea. It is named after French explorer Jean-Michel Huon de Kermadec. The peninsula is dominated by the steep Saruwaged and Finist ...
. It was originally described as a separate species in 1919 on the basis of differences in the colour of the culmen. It has an all-yellow bill in comparison to the nominate, which has a dark ridge along the culmen. It is also smaller in size. As these two subspecies are poorly defined, some authors have considered lumping the two, which would make the species
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
.


Description

It is a small kingfisher, with males weighing and females weighing . Adults are between in length. Males have a
rufous Rufous () is a color that may be described as reddish-brown or brownish- red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a d ...
head and underparts and greenish-blue upperparts, along with black loral and
nape The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , ). The corresponding adjective is ''nuchal'', as in the term ''nuchal rigidity'' ...
patches. The tail is dark blue and it has black wing feathers with bluish edges and tips. Females have a black crown and paler underparts. The neck patches are joined at the nape in females, making a bar across the back of the neck. The bill is bright yellow with a dark ridge along the culmen. The iris is dark brown and the feet and legs are dull yellow. Juveniles have a grayish-black bill and mottled-black ear coverts. They also have a larger black area around the eye and dark-tipped feathers on the cheeks and breast. The yellow-billed kingfisher is similar in appearance, but has an all-yellow bill instead and is found at lower altitudes than the mountain kingfisher. It is also larger with a proportionally larger bill.


Vocalisations

It typically vocalises from the canopy and mid-story. It has a loud whistled trilling call, typically composed of 3–4 descending trills separated by pauses. The song ends with a distinctive trill that drops, rises and drops in pitch. The initial trills are similar to the song of the chestnut-breasted cuckoo, but are longer and richer, and the cuckoo's song is much higher-pitched. The song is also similar to that of the yellow-billed kingfisher, but is deeper than that of the latter.


Distribution and habitat

The mountain kingfisher is endemic to the island of New Guinea, and is found in both
Western New Guinea Western New Guinea, also known as Papua, Indonesian New Guinea, and Indonesian Papua, is the western half of the island of New Guinea, formerly Dutch and granted to Indonesia in 1962. Given the island is alternatively named Papua, the region ...
and
Papua New Guinea Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
. It is common to uncommon through most of the
New Guinea Highlands The New Guinea Highlands, also known as the Central Range or Central Cordillera, is a long chain of mountain ranges on the island of New Guinea, including the island's highest peak, Puncak Jaya, Indonesia, , the highest mountain in Oceania. The r ...
, except for the
Bird's Head The Bird's Head Peninsula ( Indonesian: , , meaning Bird's Head in Indonesian and Dutch) or Doberai Peninsula (''Semenanjung Doberai'') is a large peninsula that makes up the northwest portion of the island of New Guinea, comprising the Indones ...
peninsula. Reports of the species from the Adelbert Mountains and the
Foja Mountains The Foja Mountains (Foja Range, Foya Mountains) () are located just north of the Mamberamo river basin in Papua, Indonesia. The mountains rise to , and have 3,000 square kilometers of old growth tropical rainforest in the interior part of the r ...
are unconfirmed. The species mostly occurs between elevations of , although they have been reported at elevations as low as . At lower elevations, the species is replaced by the yellow-billed kingfisher. The mountain kingfisher mainly inhabits
primary forest An old-growth forest or primary forest is a forest that has developed over a long period of time without Disturbance (ecology), disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organizati ...
and
secondary growth In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips ...
within its range.


Behaviour and ecology

It feeds on small
lizards Lizard is the common name used for all squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The ...
,
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s, and
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e. It hunts by perching in middle and upper canopy for long periods of time, before diving to capture prey. Nests are dug into earthen riverbanks and dry timber. On Papua New Guinea, eggs are laid in December. Eggs are laid in clutches of two and are white in color.


Status

The mountain kingfisher's population has not been quantified. However, due to its relatively large range and stable population, the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
lists it as a
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
species. It is reported as being fairly common locally but rare generally across New Guinea.


References

{{Authority control mountain kingfisher Birds of the New Guinea Highlands mountain kingfisher Taxonomy articles created by Polbot