The Celebes warty pig (''Sus celebensis''), also called Sulawesi warty pig or Sulawesi pig, is a species in the pig genus ('' Sus'') that lives on

Sulawesi Sulawesi ( ), also known as Celebes ( ), is an island in Indonesia. One of the four Greater Sunda Islands, and the List of islands by area, world's 11th-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Min ...

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...

. It survives in most habitats and can live in altitudes of up to . It has been

domesticated Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a steady supply of reso ...

and introduced to a number of other islands in Indonesia.

Description

The Celebes warty pig is a medium-sized pig, and quite variable in size and appearance. Although a number of

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

have been recognised, it is now regarded as a

monotypic taxon In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of Genus, genera, the term ...

. It is the only pig species that has been domesticated apart from the

wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a Suidae, suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The speci ...

; being semi-domesticated may have had an influence on the variability of its appearance. This pig has a head-and-body length of between and a long tail, with males generally being larger than females. The back is rounded and the legs short. The colour is greyish-black, sometimes tinged with red or yellow on the flanks. There are three pairs of facial warts and a fringe of pale bristles on the snout and more bristles on the cheeks. The crown and nape are topped by a short crest of dark bristles, while a dark mid-dorsal stripe extends from the crest towards the tufted tail. Young pigs have longitudinal stripes, but these fade as the piglets grow.

Distribution and habitat

The Celebes warty pig occurs in Sulawesi, being plentiful in central, eastern and south-eastern parts of the island but uncommon in the northeastern and southern parts. It also occurs naturally on the nearby smaller islands of

Buton Buton (also Butung, Boeton or Button) is an island in Indonesia located off the southeast peninsula of Sulawesi. It covers roughly 4,727 square kilometers in area, or about the size of Madura; it is the 129th List of islands by area, largest is ...

, Muna, Kabeana, Peleng, Lembeh and the

Togian Islands The Togian (or Togean) Islands are an archipelago of 56 islands and many offshore islets, situated in the Gulf of Tomini, off the coast of Central Sulawesi, in Indonesia. The largest islands are Batudaka Island, Batudaka, Togean Island, Togea ...

. Besides this, it has been domesticated and introduced to various other islands, has hybridised with ''

Sus scrofa The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is n ...

'', and has become

feral A feral (; ) animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in som ...

in some places, giving rise to a number of different pig populations. It inhabits various habitat types including rainforest, swamp, cultivated land and grassland, at altitudes up to .

Ecology

This pig tends to move around in small groups of up to nine individuals, led by a dominant male and including several females and their offspring. Foraging takes place mainly in the early morning and the evening; the diet consists mainly of roots, shoots, leaves and fallen fruit, but also includes carrion,

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

s and small

vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...

s. Breeding takes place throughout the year. Gestation periods are probably between four and five months, and litter sizes can be up to eight, but in one study, averaged about two. The most important natural

predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...

is the

reticulated python The reticulated python (''Malayopython reticulatus'') is a Pythonidae, python species native to South Asia, South and Southeast Asia. It is the world's List of largest snakes, longest snake, and the list of largest snakes, third heaviest snake. I ...

Conservation status

The chief threats faced by this pig are an expanding human population, deforestation, with conversion of the land to agricultural use, and hunting for human consumption. Even in areas such as national parks, where the species is protected, hunting still occurs and the meat is traded in local markets. This over-hunting and

loss of habitat Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...

has led the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...

to assess the pig's conservation status as being "

near threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...

Cave art

cave painting In archaeology, cave paintings are a type of parietal art (which category also includes petroglyphs, or engravings), found on the wall or ceilings of caves. The term usually implies prehistoric art, prehistoric origin. These paintings were often c ...

of a Celebes warty pig in the Leang Tedongnge cave on Sulawesi has been dated to an age of at least 45,500 years ago, making it the oldest known representational cave painting. A layer of

calcite Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on ...

over the painting was dated using

uranium Uranium is a chemical element; it has chemical symbol, symbol U and atomic number 92. It is a silvery-grey metal in the actinide series of the periodic table. A uranium atom has 92 protons and 92 electrons, of which 6 are valence electrons. Ura ...

-series isotope dating to give a minimum age for the painting. The image was made using

red ochre Ochre ( ; , ), iron ochre, or ocher in American English, is a natural clay earth pigment, a mixture of ferric oxide and varying amounts of clay and sand. It ranges in colour from yellow to deep orange or brown. It is also the name of the col ...

, and is accompanied by images of human hands made by blowing ochre on to the cave wall around the hands. The painting depicts a male pig, including the male's pair of facial warts. It was first seen by Westerners in 2017.

Geology

At one point in time there may have been a portion of the land which shifted upward resulted in increased diversification. The researchers found out that the Sulawesi who roam the island today were preceded by other ancestors who once expanded from the innermost area of the island to the outermost part of it.

References

Frantz, L., Rudzinski, A., Nugraha, A., Evin, A., Burton, J., Hulme-Beaman, A., Linderholm, A., Barnett, R., Vega, R., Irving-Pease, E. K., Haile, J., Allen, R., Leus, K., Shephard, J., Hillyer, M., Gillemot, S., van den Hurk, J., Ogle, S., Atofanei, C., Thomas, M. G., ... Larson, G. (2018). Synchronous diversification of Sulawesi's iconic artiodactyls driven by recent geological events. Proceedings. Biological sciences, 285(1876), 20172566. https://doi.org/10.1098/rspb.2017.2566

External links

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The Ultimate Ungulate's entry on Sus celebensis
{{Taxonbar, from=Q535406 Sus (genus) Mammals of Sulawesi Endemic fauna of Indonesia Mammals described in 1843 Taxa named by Salomon Müller Taxa named by Hermann Schlegel