Surcula Faxensis
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''Turricula'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...
s, marine
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
s in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Clavatulidae Clavatulidae is a Taxonomy (biology), taxonomic family (biology), family of sea snails, marine gastropod mollusks in the superfamily Conoidea. The family is not well differentiated morphologically. Clavatulidae was raised, based on cladistic an ...
.


Taxonomy

Clavatulidae was raised, based on
cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
analysis, from subfamily to the family level by Rosenberg in 1998. It remained regarded as a subfamily of
Turridae Turridae is a Taxonomy (biology), taxonomic family (biology), family name for a number of predatory sea snails, Marine (ocean), marine gastropod mollusks in the superfamily Conoidea.MolluscaBase (2018). Turridae H. Adams & A. Adams, 1853 (1838). ...
by several malacologists (Kantor, Sysoev).World Register of Marine Species: Clavatulidae
accessed 1 April 2010.
until a new publication in 2011


Fossil records

This genus is known in the fossil records from the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
to the
Quaternary The Quaternary ( ) is the current and most recent of the three periods of the Cenozoic Era in the geologic time scale of the International Commission on Stratigraphy (ICS), as well as the current and most recent of the twelve periods of the ...
(age range: from 70.6 to 0.0 million years ago). Fossils of species within this genus have been found all over the world.Fossilworks
/ref>


Description

The turriculated shell is fusiform with a long
spire A spire is a tall, slender, pointed structure on top of a roof of a building or tower, especially at the summit of church steeples. A spire may have a square, circular, or polygonal plan, with a roughly conical or pyramidal shape. Spire ...
. The anal sinus is situated in the infrasutural depression above the peripheral carina. The
siphonal canal The siphonal canal is an anatomical feature of the shells of certain groups of sea snails within the clade Neogastropoda. Some sea marine gastropods have a soft tubular anterior extension of the mantle called a siphon through which water i ...
is long, tapering and slightly bent. The operculum has a medio-lateral nucleus. The eyes of the animal are situated at the base of the tentacles.
Radula The radula (; : radulae or radulas) is an anatomical structure used by mollusks for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters ...
formula : 1-0-1.


Distribution

The species of this marine genus occurs in the Indo-West Pacific, from the
Red Sea The Red Sea is a sea inlet of the Indian Ocean, lying between Africa and Asia. Its connection to the ocean is in the south, through the Bab-el-Mandeb Strait and the Gulf of Aden. To its north lie the Sinai Peninsula, the Gulf of Aqaba, and th ...
and east Africa to Australia (
Northern Territory The Northern Territory (abbreviated as NT; known formally as the Northern Territory of Australia and informally as the Territory) is an states and territories of Australia, Australian internal territory in the central and central-northern regi ...
,
Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...
,
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
).


Species

Species within the genus ''Turricula'' include: * ''
Turricula aethiopica ''Turricula aethiopica'' is a species of sea snail, a marine gastropod mollusk in the family Clavatulidae. Description Distribution This species occurs in the Indian Ocean off Somalia and Madagascar Madagascar, officially the Republi ...
'' (Thiele, 1925) * ''
Turricula amplisulcus ''Turricula amplisulcus'' is a species of sea snail, a marine gastropod mollusk in the family Clavatulidae. Description Distribution This marine species occurs along Table Bay and Agulhas Bank The Agulhas Bank (, from Portuguese for Cape A ...
'' (Barnard, 1958) * † ''
Turricula bantamensis ''Turricula'', turret Turret may refer to: * Turret (architecture), a small tower that projects above the wall of a building * Gun turret, a mechanism of a projectile-firing weapon * Optical microscope#Objective turret (revolver or revolving nose ...
'' (K. Martin, 1895) * † ''
Turricula barreti ''Turricula barreti'' is an extinct species of sea snail, a marine gastropod mollusk in the family Clavatulidae.MolluscaBase eds. (2023). MolluscaBase. Turricula barreti (de Boury, 1899) †. Accessed through: World Register of Marine Species at ...
'' (de Boury, 1899) * † ''
Turricula beamensis ''Turricula'', turret Turret may refer to: * Turret (architecture), a small tower that projects above the wall of a building * Gun turret, a mechanism of a projectile-firing weapon * Optical microscope#Objective turret (revolver or revolving nose ...
'' M.A. Peyrot, 1925 * † ''
Turricula belgica ''Turricula'', turret Turret may refer to: * Turret (architecture), a small tower that projects above the wall of a building * Gun turret, a mechanism of a projectile-firing weapon * Optical microscope#Objective turret (revolver or revolving nose ...
'' (G.G. Münster, 1826) * † ''
Turricula beyrichi ''Turricula'', turret Turret may refer to: * Turret (architecture), a small tower that projects above the wall of a building * Gun turret, a mechanism of a projectile-firing weapon * Optical microscope#Objective turret (revolver or revolving nose ...
'' (R.A. Philippi, 1846) * † ''
Turricula brevicauda ''Turricula brevicauda'' is an extinct species of sea snail, a marine gastropod mollusk in the family Clavatulidae.MolluscaBase eds. (2023). MolluscaBase. Turricula brevicauda (Deshayes, 1834) †. Accessed through: World Register of Marine Spec ...
'' (Deshayes, 1834) * '' Turricula catena'' (Reeve, 1843) * † '' Turricula catenata'' (Lamarck, 1804) * † '' Turricula ceylonica'' (Smith E. A., 1877) * '' Turricula crassicosta'' (F. E. Edwards, 1857) † * † '' Turricula dentata'' (Lamarck, 1804) * † '' Turricula drilliaeformis'' (K. Martin, 1895) * † '' Turricula eastoni'' (K. Martin, 1914) * † '' Turricula everwyni'' (K. Martin, 1884) * '' Turricula faurei'' (Barnard, 1958) * † '' Turricula gembacana'' (K. Martin, 1884) * '' Turricula gemmulaeformis'' (Thiele, 1925) * † '' Turricula glyphana'' (Bayan, 1873) * '' Turricula granobalteus'' (Hedley, 1922) * † '' Turricula guttata'' (K. Martin, 1935) * † '' Turricula hillegondae'' (K. Martin, 1914) * † '' Turricula ijzermani'' (K. Martin, 1933) * † '' Turricula inermiformis'' (Csepreghy-Meznerics, 1969) * '' Turricula javana'' (Linnaeus, 1767) * † '' Turricula jogjacartensis'' (K. Martin, 1931) * † '' Turricula kelirensis'' (K. Martin, 1916) * † '' Turricula lepidota'' (K. Martin, 1914) * † '' Turricula lirocostata'' Cossmann, 1899 * † '' Turricula mauritii'' (de Boury, 1899) * † '' Turricula michelini'' (Deshayes, 1865) * '' Turricula navarchus'' (Melvill & Standen, 1903) * '' Turricula nelliae'' (Smith E. A., 1877) * † '' Turricula orthocolpa'' Cossmann, 1913 * '' Turricula panthea'' Dall, 1919 * † '' Turricula plagiaria'' (K. Martin, 1931) * † '' Turricula plateaui'' (Cossmann, 1889) * † '' Turricula pollii'' (Icke & K. Martin, 1907) * † '' Turricula polycesta'' (Bayan, 1873) * '' Turricula profundorum'' (Smith E. A., 1896) * † '' Turricula rembangensis'' (K. Martin, 1906) * † '' Turricula rostrata'' Solander in Brander, 1766 (deleted in GBIF) * † '' Turricula samarangana'' (K. Martin, 1884) * † '' Turricula sanctistephani'' Lozouet, 2017 * † '' Turricula smithi'' (K. Martin, 1884) * † '' Turricula streptopleura'' (Cossmann, 1900) * † '' Turricula subelegans'' (d'Orbigny, 1850) * '' Turricula sulcicancellata'' (Barnard, 1958) * † '' Turricula sultani'' (K. Martin, 1914) * '' Turricula sumatrana'' (Thiele, 1925) * † '' Turricula terae'' Oostingh, 1938 † * † '' Turricula textiliosa'' (Deshayes, 1834) * '' Turricula thurstoni'' (Smith E. A., 1896) * '' Turricula tornata'' (Dillwyn, 1817) * † '' Turricula transversaria'' (Lamarck, 1804) * '' Turricula turriplana'' (Sowerby III, 1903) * † '' Turricula vaudini'' (Deshayes, 1865) † * † '' Turricula veslensis'' (Cossmann, 1899) † * † '' Turricula vibekeae'' Schnetler & M. S. Nielsen, 2018 * † '' Turricula wanneri'' (Tesch, 1915) * † '' Turricula waringinensis'' (K. Martin, 1895) ;Extinct: * † '' Turricula dimidiata'' Brocchi, 1814Fossile
/ref> * † '' Turricula eolavinia'' Olsson, 1930 * † '' Turricula excelsa'' Bohm, 1926 * † '' Turricula hanguensis'' Cox, 1930 * † '' Turricula haydeni'' Cox, 1930 * † '' Turricula kayalensis'' Dey, 1961 * † '' Turricula promensis silistrensis'' Vredenburg, 1921 * † '' Turricula promensis'' Vredenburg, 1921 * † '' Turricula sethuramae'' Vredenburg, 1921 * † '' Turricula spuria'' Hedley, 1922 * † '' Turricula taurina'' Olsson, 1922 * † '' Turricula terryi'' Olsson, 1922 * † '' Turricula thangaensis'' Vredenburg, 1921


Synonyms

* ''Turricula bijubata'' (Reeve, 1843): synonym of ''
Turridrupa bijubata ''Turridrupa bijubata'', common name the crested turrid, is a species of sea snail, a marine gastropod mollusk in the family (biology), family Clavatulidae. Description The size of an adult shell varies between 11 mm and 25 mm. The c ...
'' (Reeve, 1843) * † ''Turricula (Crenaturricula) bouryi'' Glibert, 1960: synonym of † ''
Turricula barreti ''Turricula barreti'' is an extinct species of sea snail, a marine gastropod mollusk in the family Clavatulidae.MolluscaBase eds. (2023). MolluscaBase. Turricula barreti (de Boury, 1899) †. Accessed through: World Register of Marine Species at ...
'' (de Boury, 1899) (unnecessary replacement name for ''Pleurotoma barreti'' de Boury, 1899) * ''Turricula brunonia'' Dall, 1924: synonym of '' Veprecula brunonia'' (Dall, 1924) * ''Turricula cadaverosa'' (Reeve, 1844): synonym of '' Vexillum cadaverosum'' (Reeve, 1844) * ''Turricula casta'' H. Adams, 1872: synonym of '' Vexillum castum'' (H. Adams, 1872) * † ''Turricula cruziana'' Olsson, 1932 accepted as † '' Cruziturricula cruziana'' (Olsson, 1932) (original combination) * † ''Turricula curtata'' Marwick, 1926 accepted as † '' Marshallena curtata'' (Marwick, 1926) * ''Turricula ensyuensis'' Shikama, 1977: synonym of '' Comitas ensyuensis'' (Shikama, 1977) * † ''Turricula esdailei'' Marwick, 1926 accepted as † '' Tahuia esdailei'' (Marwick, 1926) * † ''Turricula (Fusimitra) extranea'' (Deshayes, 1865): synonym of † '' Vexillum extraneum'' (Deshayes, 1865) * ''Turricula flammea'' Schumacher, 1817: synonym of ''Turricula javana'' (Linnaeus, 1767) * † ''Turricula gendinganensis'' K. Martin, 1906 accepted as '' Unedogemmula indica'' (Röding, 1798) * ''Turricula granobalteatus'' ic synonym of ''Turricula granobalteus'' (Hedley, 1922) * ''Turricula hondoana'' Dall, 1925: synonym of '' Aforia circinata'' (Dall, 1873) * † ''Turricula ilonae'' Báldi, 1966 accepted as † '' Domenginella ilonae'' (Báldi, 1966) * ''Turricula japonica'' Dall, 1925 accepted as '' Aforia circinata'' (Dall, 1873) * ''Turricula kamakurana'' (Pilsbry, 1895): synonym of '' Comitas kamakurana'' (Pilsbry, 1895) * ''Turricula kuroharai'' Oyama, 1962: synonym of '' Comitas kuroharai'' (Oyama, 1962) * † ''Turricula lamarckii'' (Michelotti, 1847) accepted as † '' Kantoria lamarckii'' (Michelotti, 1847) ( superseded combination) * ''Turricula lavinia'' Dall, 1919: synonym of '' Ptychobela lavinia'' (Dall, 1919) * ''Turricula lavinoides'' Olsson, 1922: synonym of '' Fusiturricula lavinoides'' (Olsson, 1922) * ''Turricula libya'' Dall, 1919: synonym of '' Tiariturris libya'' (Dall, 1919) * ''Turricula limonensis'' Olsson, 1922: synonym of '' Fusiturricula lavinoides'' (Olsson, 1922) * † ''Turricula mertoni'' (K. Martin, 1914) accepted as † '' Pleurofusia mertoni'' (K. Martin, 1914) * ''Turricula modesta'' Pease, 1868: synonym of '' Vexillum gruneri'' (Reeve, 1844) · * † ''Turricula mordax'' (K. Martin, 1914) accepted as † '' Pleurofusia mordax'' (K. Martin, 1914) * ''Turricula murrawolga'' Garrard, 1961: synonym of '' Comitas murrawolga'' (Garrard, 1961) * † ''Turricula (Costellaria) paucicostata'' (Tate, 1889): synonym of † ''Austromitra lacertosa'' (Cernohorsky, 1970) * † ''Turricula permodesta'' (K. Martin, 1914) accepted as † ''Pentagoniturricula permodesta'' (K. Martin, 1914) * † ''Turricula planata'' F. W. Hutton, 1885 accepted as '' Austromitra planata'' (F. W. Hutton, 1885) * ''Turricula rectilateralis'' (G.B. Sowerby II, 1874): synonym of '' Vexillum suluense'' (A. Adams & Reeve, 1850) * ''Turricula scalaria'' (Barnard, 1958): synonym of '' Makiyamaia scalaria'' (Barnard, 1958) * † ''Turricula (Costellaria) tatei'' Cossmann, 1899: synonym of † '' Austromitra ralphi'' (Cossmann, 1900) * † ''Turricula taurina'' Olsson, 1922accepted as † '' Fusiturricula taurina'' (Olsson, 1922) * ''Turricula tornatus'' ic synonym of ''Turricula tornata'' (Dillwyn, 1817) * ''Turricula tuberculata'' (Gray, 1839): synonym of ''Turricula nelliae spuria'' (Hedley, 1922) * † ''Turricula waitaraensis'' Marwick, 1926: synonym of † '' Thatcheria waitaraensis'' (Marwick, 1926)


Taxa inquirenda

Further investigation needed: * ''Surcula aditus'' Barnard, 1969 * ''Surcula bouvieri'' Jousseaume, 1898 * ''Surcula urnula'' Thiele, 1925


References

* Li, BQ. & Li, XZ. 2008. ''Report on the two subfamilies Clavatulinae and Cochlespirinae (Mollusca : Neogastropoda : Turridae) from the China seas''. Zootaxa 1771: 31–42 * Wilson, B. 1994. ''Australian Marine Shells. Prosobranch Gastropods.'' Kallaroo, WA : Odyssey Publishing Vol. 2 370 pp.


External links


Schumacher, C. F. (1817). Essai d'un nouveau système des habitations des vers testacés. Schultz, Copenghagen. iv + 288 pp., 22 pls

Adams, H. & Adams, A. (1853–1858). The genera of Recent Mollusca; arranged according to their organization. London, van Voorst. Vol. 1: xl + 484 pp.; vol. 2: 661 pp.; vol. 3: 138 pls

Bouchet, P.; Kantor, Y. I.; Sysoev, A.; Puillandre, N. (2011). A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies. 77(3): 273–308



Nomenclaturor Zoologicus info
{{Taxonbar, from=Q3294028 Extant Maastrichtian first appearances