Stygimys Jepseni
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''Stygimys'' is an extinct
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
from the Upper
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
and
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
of
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. It was a member of the
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
order
Multituberculata Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct Order (biology), order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the M ...
within the
suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
Cimolodonta Cimolodonta is a clade of Multituberculata, multituberculate mammals that lived from the Cretaceous to the Eocene. They probably lived something of a rodent-like existence until their ecological niche was assumed by true rodents. The more basal ...
, family
Eucosmodontidae Eucosmodontidae is a poorly preserved family of fossil mammals within the extinct order Multituberculata. Representatives are known from strata dating from the Upper Cretaceous through the Lower Eocene of North America, as well as the Paleocene t ...
. The genus ''Stygimys'' ("Styx mouse") was named by R.E. Sloan and
Leigh Van Valen Leigh Van Valen (August 12, 1935 – October 16, 2010) was an American evolutionary biologist. At the time of his death, he was professor emeritus in the Department of Ecology and Evolution at the University of Chicago. Research and interests A ...
in
1965 Events January–February * January 14 – The First Minister of Northern Ireland and the Taoiseach of the Republic of Ireland meet for the first time in 43 years. * January 20 ** Lyndon B. Johnson is Second inauguration of Lynd ...
. The name comes from the
Styx In Greek mythology, Styx (; ; lit. "Shuddering"), also called the River Styx, is a goddess and one of the rivers of the Greek Underworld. Her parents were the Titans Oceanus and Tethys, and she was the wife of the Titan Pallas and the moth ...
(river of hell) from
Greek mythology Greek mythology is the body of myths originally told by the Ancient Greece, ancient Greeks, and a genre of ancient Greek folklore, today absorbed alongside Roman mythology into the broader designation of classical mythology. These stories conc ...
. The genus has also been known as ''
Catopsalis ''Catopsalis'' is a genus of extinct mammal from the Paleocene of North America. This animal was a relatively large member of the extinct order of Multituberculata. Most multituberculates were much smaller. At one time, the genus was also formal ...
'' (partly); ''
Cimexomys ''Cimexomys'' is an extinct North American mammal that lived from the Upper Cretaceous to the Paleocene. For a while, it shared the world with dinosaurs, but outlived them. It was a member of the extinct order Multituberculata and lies within the ...
'' (partly); '' Eucosmodon'' (partly); and ''
Parectypodus ''Parectypodus'' (meaning "besides ''Ectypodus''") is an extinct genus of mammals that lived from Late Cretaceous (Maastrichtian) to Eocene time in North America. It is a member of the extinct order of Multituberculata, suborder Cimolodonta, fami ...
'' (partly). Some skull material is known, but not much.


Species

The species ''Stygimys camptorhiza'' was named by P.A. Johnston and R.C. Fox in
1984 Events January * January 1 – The Bornean Sultanate of Brunei gains full independence from the United Kingdom, having become a British protectorate in 1888. * January 7 – Brunei becomes the sixth member of the Association of Southeas ...
. Place:
Puercan The Puercan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), spanning the interval from 66,000,000 to 63,300,000 years BP lasting . It is usua ...
(Paleocene)-age
strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
of Rav W-1 in
Saskatchewan Saskatchewan is a Provinces and territories of Canada, province in Western Canada. It is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and to the south by the ...
, Canada. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
for this species is in the
University of Alberta The University of Alberta (also known as U of A or UAlberta, ) is a public research university located in Edmonton, Alberta, Canada. It was founded in 1908 by Alexander Cameron Rutherford, the first premier of Alberta, and Henry Marshall Tory, t ...
collection. The species ''Stygimys cupressus'' was named by R.C. Fox in
1989 1989 was a turning point in political history with the "Revolutions of 1989" which ended communism in Eastern Bloc of Europe, starting in Poland and Hungary, with experiments in power-sharing coming to a head with the opening of the Berlin W ...
. Remains are known from the Puercan (Paleocene)-age strata of the Long Fall Horizon of Canada. The species ''Stygimys jepseni'' was named by
George Gaylord Simpson George Gaylord Simpson (June 16, 1902 – October 6, 1984) was an American paleontologist. Simpson was perhaps the most influential paleontologist of the twentieth century, and a major participant in the modern synthesis, contributing '' Tempo ...
in
1935 Events January * January 7 – Italian premier Benito Mussolini and French Foreign Minister Pierre Laval conclude an agreement, in which each power agrees not to oppose the other's colonial claims. * January 12 – Amelia Earhart ...
and Sloan and Van Valen in 1965. It is also known as ''Eucosmodon sparsus'' (Simpson 1937a); ''E. jepseni''; and ''Parectypodus jepseni'' (Simpson 1935d). Remains are known from the Torrejonian (Paleocene)-age strata of
Montana Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
(USA). This species has been cited as a descendant of ''S. kuszmauli''. The animal weighed about 90 g. The species ''Stygimys kuszmauli'' was named by R.E. Sloan and Leigh Van Valen in 1965. It has also been known as ''Catopsalis foliates'' (Cope 1882); ''Cimexomys gratus'' (Jepsen GL,
1930 Events January * January 15 – The Moon moves into its nearest point to Earth, called perigee, at the same time as its fullest phase of the Lunar Cycle. This is the closest moon distance at in recent history, and the next one will be on J ...
); ''Eucosmodon gratus'' (G.L. Jepson 1930); ''Eucosmodon kuszmauli ''; and ''Stygimys gratus''. Remains were found in
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
(Upper Cretaceous) - Puercan (Paleocene)-age strata of the
Polecat Bench Formation The Polecat Bench Formation is a geologic formation in Montana. It preserves fossils dating back to the Paleogene period.D. E. Russell. 1967. Le Paleocene continental d'Amerique du nord. Memoires du Museum National d'Histoire Naturelle. Serie C. ...
of
Wyoming Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
and the
Hell Creek Formation The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation s ...
of Montana. Some scraps of jaw are known. This species is the most basal member of the genus, with a weight of 300 g. The species ''Stygimys teilhardi'' was named by W. Granger & George Gaylord Simpson in
1929 This year marked the end of a period known in American history as the Roaring Twenties after the Wall Street Crash of 1929 ushered in a worldwide Great Depression. In the Americas, an agreement was brokered to end the Cristero War, a Catholic ...
and R.E. Sloan and Leigh Van Valen in 1965. It has also been known as ''Eucosmodon teilhardi'' (Granger W. & Simpson G.G. 1929). Remains are known from the
Torrejonian The Torrejonian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 63,300,000 to 60,200,000 years BP lasting . It is usually ...
(Paleocene)-age strata of the
San Juan Basin The San Juan Basin is a geologic structural basin located near the Four Corners region of the Southwestern United States. The basin covers 7,500 square miles and resides in northwestern New Mexico, southwestern Colorado, and parts of Utah a ...
of
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
. This species has been cited as a possible descendant of ''S. kuszmauli''.


References

* (Most of the information in this article has been derived from this page.) * * * * * {{Taxonbar, from=Q7629326 Cimolodonta Cretaceous mammals Paleocene mammals Paleocene genus extinctions Cretaceous mammals of North America Hell Creek fauna Prehistoric mammal genera