The stump-tailed macaque (''Macaca arctoides''), also called the bear macaque, is a species of
macaque
The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and Europe (in Gibraltar). Macaques are principally f ...
native to
South Asia
South Asia is the southern Subregion#Asia, subregion of Asia that is defined in both geographical and Ethnicity, ethnic-Culture, cultural terms. South Asia, with a population of 2.04 billion, contains a quarter (25%) of the world's populatio ...
and
Southeast Asia
Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
. In India, it occurs south of the
Brahmaputra River
The Brahmaputra is a trans-boundary river which flows through Southwestern China, Northeastern India, and Bangladesh. It is known as Brahmaputra or Luit in Assamese language, Assamese, Yarlung Tsangpo in Lhasa Tibetan, Tibetan, the Siang/Dihan ...
, in the northeastern part of the country. Its range in India extends from
Assam
Assam (, , ) is a state in Northeast India, northeastern India, south of the eastern Himalayas along the Brahmaputra Valley, Brahmaputra and Barak River valleys. Assam covers an area of . It is the second largest state in Northeast India, nor ...
and
Meghalaya
Meghalaya (; "the abode of clouds") is a states and union territories of India, state in northeast India. Its capital is Shillong. Meghalaya was formed on 21 January 1972 by carving out two districts from the Assam: the United Khasi Hills an ...
to eastern Arunachal Pradesh, Nagaland, Manipur, Mizoram and Tripura.
It is primarily
frugivorous
A frugivore ( ) is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance ...
but eats many types of vegetation, such as seeds, leaves and roots. It also hunts
freshwater crab
Around 1,300 species of freshwater crabs are distributed throughout the tropics and subtropics, divided among eight family (biology), families. They show direct development and maternal care of a small number of offspring, in contrast to marine c ...
s,
frog
A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s,
bird egg
Bird eggs are laid by the females and range in quantity from one (as in condors) to up to seventeen (the grey partridge). Avian clutch size, Clutch size may vary latitudinally within a species. Some birds lay eggs even when the eggs have not been ...
s and
insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s.
Characteristics
The stump-tailed macaque has long, thick, dark brown fur covering its body, but its face and its short tail, which measures between , are hairless.
[ Infants are born white and darken as they mature.][ As they age, their bright pink or red faces darken to brown or nearly black and lose most of their hair.][ Males are larger than females, measuring long and weighing , while females measure and weigh .][ Males' ]canine teeth
In mammalian oral anatomy, the canine teeth, also called cuspids, dogteeth, eye teeth, vampire teeth, or fangs, are the relatively long, pointed teeth. In the context of the upper jaw, they are also known as '' fangs''. They can appear more fl ...
, which are important for establishing dominance within social groups, are more elongated than those of the females.[ Like all macaques, this species has cheek pouches to store food for short periods of time.][
]
Distribution and habitat
This Old World monkey
Old World monkeys are primates in the family Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons (genus '' Papio''), red colobus (genus '' Piliocolob ...
travels quadrupedally, usually on the ground, as it is not very agile in trees.[ It is generally found in ]evergreen
In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
tropical and subtropical moist broadleaf forests
Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF).
Description
TSMF is generally found in la ...
, at different elevation
The elevation of a geographic location (geography), ''location'' is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational equipotenti ...
s depending on the amount of rainfall in the area.[ It depends on ]rainforest
Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s for food and shelter, and is not found in dry forests except where it ranges in the Himalayan region of India, only spending time in secondary forest
A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
s if it is bordering old-growth forest
An old-growth forest or primary forest is a forest that has developed over a long period of time without disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organization of the United Natio ...
tropical forests.[ With its thick fur, the stump-tailed macaque can live in cold climates at elevations up to . It is distributed from northeastern ]India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
and southern China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
into the northwest tip of West Malaysia
Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
on the Malay Peninsula
The Malay Peninsula is located in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area contains Peninsular Malaysia, Southern Tha ...
.[ It is also found in ]Burma
Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and ha ...
, Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
, Laos
Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...
, Cambodia
Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
and Vietnam
Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
.[ It is possibly extinct in ]Bangladesh
Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...
. No global population estimate is available. In Cambodia, a declining population of 230 is reported from Keo Seima Wildlife Sanctuary
Keo Seima Wildlife Sanctuary (Khmer language, Khmer: ដែនជម្រកសត្វព្រៃកែវសីមា) is a protected area of mixed seasonal tropical forest in eastern Cambodia, located in Mondulkiri Province, Mondulkiri and ...
.
A study population was introduced to Tanaxpillo, an uninhabited island in Lake Catemaco, Veracruz
Veracruz, formally Veracruz de Ignacio de la Llave, officially the Free and Sovereign State of Veracruz de Ignacio de la Llave, is one of the 31 states which, along with Mexico City, comprise the 32 Political divisions of Mexico, Federal Entit ...
, Mexico
Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
in 1974, where it ranges in seminatural conditions.[ Most information on the species comes from the introduced population on Tanaxpillo and other captive settings, as few long-term studies have been conducted on the stump-tailed macaque in the wild.][
]
Behaviour and ecology
Stump-tailed macaque generally share the same social structure of any macaque species, with a linear, hereditary dominance hierarchy hereditary in females yet fluctuates among males based on their fighting ability and social maneuvering. Compared to rhesus and long-tail macaques, stump-tails defuse intense confrontations, are relatively tolerant of group members, and have a surprisingly rich repertoire of reconciliation tactics. This species lacks pair bonds and is promiscuous, a staple for macaques. Physical violence very rarely occurs, and although minor scraps often flair up and physical intimidation displays occur, they tend to quickly be resolved, resulting in this species being labelled as peaceful. Stump-tailed macaques are remarkably unfussy in their eating habits, though fruit generally is a staple part of their diet. Stump-tail macaques have a large, bulky, muscular build with thick, solid limbs, making them very mobile on land yet quite ungainly in trees, and this unusual physique for a macaque may be responsible for this species unique tendency to consume larger quantities of meat than other macaque species. Stump-tail macaques feed on very large quantities of insects, small animals and eggs.
Reproduction
A study population of female stump-tailed macaques was found to have increased levels of steroid sex hormones, specifically 17β-estradiol and progesterone levels. 17β-estradiol levels were significantly greater during summer and fall and progesterone levels were significantly greater during summer, fall and winter. This explains how stump-tailed macaques have two mating seasons per year: one in summer (July–August) and one in fall (November). This is supported by the distribution of birth frequency in stump-tailed macaques.
See also
* Britches – an infant stump-tailed macaque used in sight-deprivation experiments, which was stolen from the laboratory by the Animal Liberation Front
The Animal Liberation Front (ALF) is a Far-left politics, far-left international, Leaderless resistance, leaderless, decentralized movement that emerged in Britain in the 1970s, evolving from the Bands of Mercy. It operates without a formal lead ...
.
* List of endangered and protected species of China
A list is a set of discrete items of information collected and set forth in some format for utility, entertainment, or other purposes. A list may be memorialized in any number of ways, including existing only in the mind of the list-maker, bu ...
References
{{Authority control
stump-tailed macaque
Primates of Southeast Asia
Mammals of Bangladesh
Mammals of India
Mammals of Thailand
Mammals of Vietnam
Mammals of Laos
Mammals of Cambodia
Mammals of Malaysia
Mammals of China
Mammals of Myanmar
Fauna of Yunnan
stump-tailed macaque
Taxa named by Isidore Geoffroy Saint-Hilaire