Struthiola Striata
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''Struthiola striata'' is a rounded, heather-like shrub of up to high that is assigned to the
Thymelaeaceae The Thymelaeaceae are a cosmopolitan family (biology), family of flowering plants composed of 50 genera (listed below) and 898 species.Zachary S. Rogers (2009 onwards)A World Checklist of Thymelaeaceae (version 1) Missouri Botanical Garden Webs ...
family. It has small assending leaves on long straight branches, with cream, soft yellow or pinkish flowers in spikes, each of which consist of a tube of about long with 4 oval sepal lobes and 4 yellow alternating petal-like scales. It is sometimes called ribbed capespray or featherhead in English and roemenaggie, katstertjie or veërtjie in
Afrikaans Afrikaans is a West Germanic languages, West Germanic language spoken in South Africa, Namibia and to a lesser extent Botswana, Zambia, Zimbabwe and also Argentina where there is a group in Sarmiento, Chubut, Sarmiento that speaks the Pat ...
. It grows on coastal flats and foothills in the Western Cape province of South Africa.


Taxonomy

This species was first described as ''Struthiola striata'' by
Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biologi ...
in 1792, based on a collection by the French gardener-botanist Joseph Martin in the Cape area (Cap B. Spei). Later botanists however, overlooked this name. Franz Wilibald Schmidt named the species ''S. sulcata'' in 1793,
Henry Cranke Andrews Henry Cranke Andrews (c. 1759 – 1835, floruit, fl. 1794 – 1830), was an English botanist, botanical artist and engraver. As he always published as Henry C. Andrews, and due to difficulty finding records, the C. was often referred to as Cha ...
in 1800 created the name ''S. imbricata'', while
Jens Wilken Hornemann Jens Wilken Hornemann (6 March 1770 – 30 July 1841) was a Danish botanist. Biography He was a lecturer at the University of Copenhagen Botanical Garden from 1801. After the death of Martin Vahl in 1804, the task of publishing the Flora Danica ...
assigned the name ''S. lateriflora'' to it in 1815. In 1843 Meissner distinguished two varieties, ''S. striata'' var. ''striata'' and ''S. striata'' var. ''angustifolia''. In 1857 he considered ''S. sulcata'', ''S. imbricata'' and ''S. lateriflora''
synonyms A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
of ''S. striata''.
Charles Henry Wright Charles is a masculine given name predominantly found in English and French speaking countries. It is from the French form ''Charles'' of the Proto-Germanic name (in runic alphabet) or ''*karilaz'' (in Latin alphabet), whose meaning was ...
in 1915 in turn regarded ''S. striata'' var. ''angustifolia'' a synonym of ''S. striata''.


Description

Ribbed capespray is a rounded shrub of up to high that emerges from a single trunk from the ground. The branches are initially covered in soft hairs pressed firmly to the surface, but become hairless when aging. The leaves are in pairs opposing each other, more or less upright, overlapping each other, with the blade directly attached to the branche without a leaf stalk. They are oval to lance-shaped, long and generally wide, somewhat depressed along the main vein, almost pointed, with a row of hairs pressed to the surface along its rim. When the leaves are young 3 to 5 stripes are visible along the length of the outward facing surface. The flowers are seated in the axils of the leaves over a considerable length towards the tip of the branches, an inflorescence type called a
spike Spike, spikes, or spiking may refer to: Arts, entertainment, and media Books * ''The Spike'' (novel), a novel by Arnaud de Borchgrave * ''The Spike'' (Broderick book), a nonfiction book by Damien Broderick * ''The Spike'', a starship in Peter ...
. The two
bracteoles In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also look ...
are set opposite each other at the base of the flower, awl-shaped, about long, with a regular row of hairs along the margin and a blunt tip ending in a tuft of hairs. The
calyx CALYX, Inc. is a non-profit publisher of art and literature by women founded in 1976 based in Corvallis, Oregon. CALYX publishes both '' CALYX, A Journal of Art and Literature by Women'' twice a year and CALYX Books, which publishes one to three ...
is cream-coloured, with at its base a cylindrical tube of about long that is covered on the outside with soft hairs pressed against its surface and at its top 4 oval protracted, almost pointy lobes of long and wide, hairless but for a tuft of hairs at the tips of the outer pair. Four yellow, elliptic, fleshy, petal-like scales are implanted above and alternating with the sepal lobes, and are surrounded by stiff hairs that may be shorter or longer than the scales. About below the mouth of the tube sits on whorl of four seated stamens, that have a reddish wart at their tips. The
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
is egg-shaped, hairless and about 1 mm long, and is topped by a
style Style, or styles may refer to: Film and television * ''Style'' (2001 film), a Hindi film starring Sharman Joshi, Riya Sen, Sahil Khan and Shilpi Mudgal * ''Style'' (2002 film), a Tamil drama film * ''Style'' (2004 film), a Burmese film * '' ...
of about long and ends in a brush-like stigma.


Differences with related species

''Struthiola'' has its flowers arranged in long spikes and each flower has 4 anthers, whereas ''
Gnidia ''Gnidia'' is a genus of flowering plants in the family Thymelaeaceae. It is distributed in tropical and southern Africa and Madagascar; more than half of all the species are endemic to South Africa.Rogers, Z. S. (2006)A new species of Malagasy ' ...
'' differs from it in having its flowers in short spikes, solitary or more often in heads, and the flowers have 8 or 10 anthers in 2 whorls. Both '' Lachnaea'', ''
Passerina The genus ''Passerina'' is a group of birds in the cardinal family (Cardinalidae). Although not closely related to the buntings in the family Emberizidae, they are sometimes known as the North American buntings. The males show vivid colors i ...
'' and a few ''Gnidia''-species lack scales on the calyx tubes. Both ''S. striata'' and '' S. tetralepis'' have four petal-like scales implanted at the opening of the calyx tube, while the many dozens of remaining ''Struthiola'' species have eight or twelve scales. ''S. striata'' is a roundish shrub of up to 1.5 m high with cream, soft yellow or pinkish flowers ending in oval sepal lobes of 1–2 mm long and 1.5–2.5 mm wide, and a reddish wart on the tip of the anthers, ''S. tetralepis'' is a single stemmed shrublet of up to high that has initially greenish yellow, later brownish red flowers with lance-shaped sepal lobes of long and wide, and a white wart on the anthers.


Distribution, ecology and conservation

''Struthiola striata'' is restricted to coast of the Western Cape province of South Africa, between
Yzerfontein Yzerfontein, or Ysterfontein, is a small harbour town with about 1200 inhabitants on the west coast of South Africa about 90 km north of Cape Town. The name in Afrikaans means "Iron Fountain".The town started out when the farm 'Yzerfontein' ...
in the north and
Mossel Bay Mossel Bay () is a harbour town of about 170,000 people on the Garden Route of South Africa. It is an important tourism and farming region of the Western Cape Province. Mossel Bay lies 400 kilometres east of the country's seat of parliament, Ca ...
in the east. It occurs in a vegetation type called sand fynbos where it grows on lower slopes and coastal sand flats. It is honey-scented at night and gets pollinated by moths. The continued survival of this species is considered to be of
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
because its populations are stable.


References

{{Taxonbar, from=Q17582202 Struthiola striata Endemic flora of the Cape Provinces