Struthiola
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''Struthiola'' is a genus of plants in the family
Thymelaeaceae The Thymelaeaceae are a cosmopolitan family (biology), family of flowering plants composed of 50 genera (listed below) and 898 species.Zachary S. Rogers (2009 onwards)A World Checklist of Thymelaeaceae (version 1) Missouri Botanical Garden Webs ...
. In habit they are
ericoid The word "ericoid" is used in modern biological terminology for its literal meanings and for extensions. Ericoid could have more than one meaning, but in practice the most common use is in reference to a plant's habit, to describe small, tough (s ...
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s or shrublets. The genus includes 31 species native to Africa, which range from
Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
to
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
.


Overview

There are forty-odd species, mainly South African, mainly occurring in the Western Cape, about 25 endemic to
fynbos Fynbos (; , ) is a small belt of natural shrubland or heathland vegetation located in the Western Cape and Eastern Cape provinces of South Africa. The area is predominantly coastal and mountainous, with a Mediterranean climate. The fynbos ...
. Their leaves are usually opposite, but sometimes alternate. Their flowers are sessile and generally solitary, but sometimes in pairs in the axils of the upper leaves. Each flower is accompanied by two ciliate bracteoles. The calyx is roughly cylindrical, with four lobes joined into a tube. The lobes are ovate to linear. There are four or eight or even twelve fleshy, subterete petals (Manning refers to them as petal-scales or petal-like-scales.) The petals are shorter than the calyx-lobes, and are surrounded by short hairs. There are four stamens arising from deep in throat of calyx-tube. The anthers are subsessile and linear, sometimes with an apical appendage. The ovary has a single loculus with a single glabrous ovule. The
style Style, or styles may refer to: Film and television * ''Style'' (2001 film), a Hindi film starring Sharman Joshi, Riya Sen, Sahil Khan and Shilpi Mudgal * ''Style'' (2002 film), a Tamil drama film * ''Style'' (2004 film), a Burmese film * '' ...
is lateral, with a simple stigma, which is usually penicillate with short hairs. The fruit is small, typically 1–3mm, dry, included in the persistent base of the calyx-tube.


Naming and etymology

The structure of the fruit in its calyx-remnant has been likened to the beak of a sparrow and gave rise to the name "''Struthiola''", from the Latin ''strutheus'' for a sparrow or ''strouthos'' for any small bird. As usual for such inconspicuous plants, common names are neither very helpful, nor consistent. The name "stroop bossie" ("syrup bush") is locally applied to ''Struthiola ciliata'', suggesting that rural children might have been experimenting with sucking the flowers, but such explanations are necessarily speculative. One or all species are referred to as "gonna", sometimes qualified, for example "soetgonna" (meaning "sweet gonna", for ''Struthiola dodecandra''), or "aandgonna" ("evening gonna", for ''Struthiola argentea''). "Gonna" however, also is variously applied to the recognisably similar genus ''
Passerina The genus ''Passerina'' is a group of birds in the cardinal family (Cardinalidae). Although not closely related to the buntings in the family Emberizidae, they are sometimes known as the North American buntings. The males show vivid colors i ...
'' and to some other genera in the Thymelaeaceae, such as ''
Dais A dais or daïs ( or , American English also but sometimes considered nonstandard)dais
in the Random House Dictionary< ...
''. So are some other variations such as "ganna", but because they are informal and local, it is not practical to be authoritative about the limits to their correct application.


Species

31 species are accepted. *'' Struthiola angustiloba'' *'' Struthiola anomala'' *'' Struthiola argentea'' *'' Struthiola bachmanniana'' *'' Struthiola cicatricosa'' *'' Struthiola ciliata'' *'' Struthiola confusa'' *'' Struthiola dodecandra'' *'' Struthiola eckloniana'' *'' Struthiola ericoides'' *'' Struthiola fasciata'' *'' Struthiola garciana'' *'' Struthiola hirsuta'' *'' Struthiola leptantha'' *'' Struthiola lineariloba'' *'' Struthiola macowanii'' *'' Struthiola martiana'' *'' Struthiola montana'' *'' Struthiola mundii'' *'' Struthiola myrsinites'' *'' Struthiola parviflora'' *'' Struthiola pondoensis'' *'' Struthiola recta'' *'' Struthiola rhodesiana'' *'' Struthiola rigida'' *'' Struthiola salteri'' *'' Struthiola scaettae'' *'' Struthiola striata'' *'' Struthiola tetralepis'' *'' Struthiola thomsonii'' *'' Struthiola tomentosa''


Horticultural significance

''Struthiola'' species are not spectacular plants; in their sparse ericoid habit they are typical of fynbos scrub, though also typically of such scrub, they are very attractive in the fine detail of their flowers and structure. Curiously, in spite of their modest appearance they have been quietly popular as garden plants, particularly among collectors, perhaps because of their delicate, unexpectedly arresting nocturnal scent. Although some species, such as ''Struthiola myrsinites'', flower quite attractively, the flowering of smaller species is easily overlooked by day, though a single plant can scent a porch on a still, warm night. This is in keeping with their narrow flowers being pollinated by small moths with fine proboscises.


References

{{Taxonbar, from=Q7625447 Thymelaeoideae Thymelaeaceae genera Flora of the Afrotropical realm Taxa described in 1767 Taxa named by Carl Linnaeus