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''Stonogobiops nematodes'', the Filament-finned prawn-goby, the Antenna goby, the high-fin goby, the red-banded goby, the high-fin red-banded goby, the striped goby, the barber-pole goby, or the black-ray Goby, is a species of marine goby native to the
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and western
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from the
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to the
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and
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.


Physical features

Adult fish can grow up to in length, with the striking pointed
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
becoming more raised and pronounced in adulthood. This elongated fin is the most obvious distinguishing feature between the black-ray goby and its close cousin, the yellow snout goby (''S. Xanthorhinica''). The fish are coloured with four diagonal brown stripes across a white body, and a distinctive yellow head. It is almost impossible for anybody less than a specialised expert in the specific field of these types of fish to discern differences between males and females of the species.


Natural environment

This goby inhabits sandy or sand-rubble bottoms adjacent to reefs at depths of from . It is one of several species that form commensal relationships with Randall's pistol shrimp (''
Alpheus randalli ''Alpheus randalli'' is a species of snapping shrimp in the family Alpheidae. It lives in the Marquesas Islands and parts of the Indian Ocean, including the Seychelles, in association with a goby of the genus ''Amblyeleotris''. The shrimp is tr ...
'').


Behaviour in the wild

This species shares a burrow with its shrimp partner. The goby has much better eyesight than the shrimp, and, as such, acts as the watchman for both of them, keeping an eye out for danger. The shrimp spends the day digging a
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
in the sand in which both live. Burrows usually measure up to one inch in diameter, and can reach up to four feet in length. The two animals maintain continuous contact, with the shrimp placing one of its antennae permanently on the goby's tail. When danger threatens, the goby will make continuous flicks of its tail, warning the shrimp there is a predator nearby, and the shrimp will remain safely in the burrow. If the danger reaches a certain level, the goby will dart into the burrow after the shrimp. At night, the goby will go into the burrow, and the shrimp will collapse the entrance to close it off. The burrow is exited the next day by the goby blasting its way out and collapsing the burrow. The shrimp then spends the next day laboriously rebuilding the entrance to the burrow. Both animals have also been known to share food with each other. When the goby catches food, it will often give a portion of it to the pistol shrimp, through a somewhat similar process of a mother bird regurgitating food to her chicks. This way, the shrimp and the goby are kept well fed. Sometimes, if the shrimp is not kept well fed, it will resort to killing the goby. In the wild, most burrows are shared by male and female goby pairs, with their respective shrimp partners, and the female goby will use this burrow as a nesting site to lay her eggs. The obvious benefits to both organisms of this
symbiotic Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
relationship make the interaction a form of mutualism
Here is an interesting example
of this. The other fish is a dartfish (genus '' Ptereleotris''). These fish are often found as unwelcome but ignored guests sharing the burrow with goby and shrimp.


Aquarium keeping


Black-ray goby in commercial trade

This goby is in high demand for home and hobby marine aquaria due to its beautiful colouration, docile nature and interesting interaction with symbiotic shrimp. This type of goby is the most common Stonogobiops species to show up in the marine trade, but is still quite rare.


Behaviour and compatibility

This fish is very docile and poses almost no threat to any other stock inhabiting a typical marine aquarium. This passiveness makes it a perfect tankmate for delicate species like sea horses or pipefish. In fact, it is in reality quite shy, and when first introduced into an aquarium, may take up to several weeks before it is bold enough to leave its hiding place, or bolt hole. While this fish can display aggression towards other tank inhabitants by opening its mouth and "yawning" at them, this is mostly show, and the goby will quickly turn tail and hide if confronted. The goby will spend most of its time hovering about two inches above its bolt hole, searching for scraps of food in the water column. If scared or startled, it will slowly retreat towards its hole. If the danger does not go away, it will dart inside at lightning speed. Mated pairs of this fish are very rare and difficult to attain. Individual males may fight if placed in a tank smaller than about 50 gallons (~200 litres).


Tank environment

For successful aquarium culture, this fish needs good sand/coral rubble cover for burrow-building and much rock cover; a reef environment is suitable. The recommended minimum tank size is 10 gallons (40 litres), however these fish mainly hide out in burrows all day, and are not active swimmers, making them candidates for smaller "pico" aquariums. (Aquariums 4 gallons and under.) The water
specific gravity Relative density, or specific gravity, is the ratio of the density (mass of a unit volume) of a substance to the density of a given reference material. Specific gravity for liquids is nearly always measured with respect to water (molecule), wa ...
should be 1.020 - 1.025, with a pH of 8.1 - 8.4; water temperature at 72 - 78 °F / 22 - 25 °C is ideal, however water temperature up to 80 °F will not harm the fish.


Care and maintenance

Small meaty foods, such as
mysid Mysida is an order of small, shrimp-like crustaceans in the malacostracan superorder Peracarida. Their common name opossum shrimps stems from the presence of a brood pouch or "marsupium" in females. The fact that the larvae are reared in thi ...
(sometimes referred to as ''mysis'' shrimp) or
brine Brine is a high-concentration solution of salt (NaCl) in water (H2O). In diverse contexts, ''brine'' may refer to the salt solutions ranging from about 3.5% (a typical concentration of seawater, on the lower end of that of solutions used for br ...
shrimp, along with flake food and algae wafers, spirulina, etc. are all happily accepted. In the wild, these gobies most often feed on
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
. The water quality must be kept reasonably high, as with all marine species. A substrate of small-grained coral sand, with larger particles mixed in (preferably four inches or deeper) is ideal for the goby/shrimp pair to make their burrow.


References


External links

* ;General information
General information on this type of goby, from About.com

Some general/taxonomic info
;Shrimp-goby interaction

* ttp://www.reefkeeping.com/issues/2004-01/hcs3/index.php Excellent general information on this goby, and the genus ''Stonogobiops'' in general, along with their mutualism with pistol shrimpbr>- An interesting study analysing daily shrimp activity cycles, the effects of goby presence on shrimp behavior, and the effects of predation on numerical density and size of gobies
{{DEFAULTSORT:Black-Ray Goby Black-ray goby Symbiosis Fish described in 1982