''Stigmaria'' is a form taxon for common fossils found in Carboniferous rocks. They represent the underground rooting structures of coal forest

lycopsid Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching s ...

trees such as '' Sigillaria'' and ''

Lepidodendron ''Lepidodendron'' is an extinct genus of primitive vascular plants belonging to the family Lepidodendraceae, part of a group of Lycopodiopsida known as scale trees or arborescent lycophytes, related to quillworts and lycopsids (club mosses). T ...

''. These swamp forest trees grew to 50 meters and were anchored by an extensive network of branching underground structures with "rootlets" attached to them. Analysis of the morphology and anatomy of these stigmarian systems suggests they were shoot-like and so they are called rhizomes or rhizophores. The stigmarian rhizomes are typically covered with a spiral pattern of circular scars where "rootlets" were attached. Since the stigmarian systems are shoot-like, these "rootlets" may be modified leaves, adapted to serve the function of roots. However, some paleontologists argue that the "rootlets" were true roots, with a complex branching structure and root hairs, comparable to the roots of the closest living relative of ''Lepidodendron'', the quillworts (genus ''

Isoetes ''Isoetes'', commonly known as the quillworts, is the only extant genus of plants in the family Isoetaceae, which is in the class of lycopods. There are currently 192 recognized species, with a cosmopolitan distribution but with the individual ...

'').

References

Prehistoric lycophytes Carboniferous plants Prehistoric plants of North America Fossils of Georgia (U.S. state) Paleozoic life of New Brunswick Paleozoic life of the Northwest Territories Paleozoic life of Nova Scotia Prehistoric lycophyte genera {{Carboniferous-plant-stub