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The gull-billed tern (''Gelochelidon nilotica''), formerly ''Sterna nilotica'', is a
tern Terns are seabirds in the family Laridae that have a worldwide distribution and are normally found near the sea, rivers, or wetlands. Terns are treated as a subgroup of the family Laridae which includes gulls and skimmers and consists o ...
in the family
Laridae Laridae is a family of seabirds in the order Charadriiformes that includes the gulls, terns, skimmers and kittiwakes. It includes around 100 species arranged into 22 genera. They are an adaptable group of mostly aerial birds found worldwide. Ta ...
. It is widely distributed and breeds in scattered localities in Europe, Asia, northwest Africa, and the Americas. The
Australian gull-billed tern The Australian tern or Australian gull-billed tern (''Gelochelidon macrotarsa'') is a tern in the family Laridae. The genus name is from Ancient Greek ''gelao'', "to laugh", and ''khelidon'', "swallow". It was previously considered conspecific w ...
was previously considered a subspecies.


Taxonomy

The gull-billed tern was formally described in 1789 by the German naturalist Johann Friedrich Gmelin in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
's ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nom ...
''. He placed it with terns in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''
Sterna ''Sterna'' is a genus of terns in the bird family Laridae. The genus used to encompass most "white" terns indiscriminately, but mtDNA sequence comparisons have recently determined that this arrangement is paraphyletic. It is now restricted to t ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Sterna nilotica''. Gmelin based his description on the "Egyptian tern" that had been described in 1785 by the English ornithologist John Latham in his book ''A General Synopsis of Birds''. Latham had in turn based his own account on that by the Swedish naturalist
Fredrik Hasselquist Fredrik Hasselquist (3 January 1722 – 9 February 1752) was a Swedish traveller and naturalist. Hasselquist was born at Törnevalla, which is two kilometers east of Linghem, Östergötland. He studied under Carl Linnaeus at Uppsala Univers ...
that was published in 1757. The gull-billed tern was moved to the resurrected genus ''
Gelochelidon ''Gelochelidon'' is a genus of terns. It was considered a monotypic genus until the Australian tern was split from the gull-billed tern. Taxonomy The genus ''Gelochelidon'' was introduced in 1830 by the German zoologist Alfred Brehm. The type s ...
'' based on a
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2005. The genus had been introduced in 1830 by the German zoologist
Alfred Brehm Alfred Edmund Brehm (; 2 February 1829 – 11 November 1884) was a German zoologist, writer, director of zoological gardens and the son of Christian Ludwig Brehm, a famous pastor and ornithologist. Through the book title '' Brehms Tierl ...
. The genus name combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
''gelaō'' meaning "to laugh" with ''khelidōn'' meaning "swallow". The specific epithet ''nilotica'' is from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
''niloticus'' meaning "of the River Nile". Five subspecies are recognised: * ''G. n. nilotica'' ( Gmelin, 1789)
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...
, found in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
,
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in t ...
through the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabia (including the Arabian Peninsula and Bahrain), Asia Minor (Asian part of Turkey except Hatay Province), East Thrace (Europ ...
& south-central
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
to western China &
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
* ''G. n. affinis'' ( Horsfield, 1821) – found in Transbaikalia to Manchuria, Japan, south and east China through
southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
to the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
,
Borneo Borneo (; id, Kalimantan) is the List of islands by area, third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java Isl ...
, Sulawesi & Sumatra * ''G. n. aranea'' (
Wilson Wilson may refer to: People *Wilson (name) ** List of people with given name Wilson ** List of people with surname Wilson * Wilson (footballer, 1927–1998), Brazilian manager and defender *Wilson (footballer, born 1984), full name Wilson Rod ...
, 1814)
– found in eastern & southern United States,
Greater Antilles The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, ...
* ''G. n. vanrossemi'' Bancroft, 1929 – found from southern
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
to northwestern Mexico * ''G. n. gronvoldi'' Mathews, 1912 – found from
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
to northeastern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...


Description

This is a fairly large and powerful tern, similar in size and general appearance to a
Sandwich tern The Sandwich tern (''Thalasseus sandvicensis'') is a tern in the family Laridae. It is very closely related to the lesser crested tern (''T. bengalensis''), Chinese crested tern (''T. bernsteini''), Cabot's tern (''T. acuflavidus''), and ele ...
, but the short thick gull-like
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
, broad wings, long legs and robust body are distinctive. The summer adult has grey upperparts, white underparts, a black cap, strong black bill and black legs. The call is a characteristic ''ker-wik''. It is in length and in wingspan. Body mass ranges from . In winter, the cap is lost, and there is a dark patch through the eye like a
Forster's tern Forster's tern (''Sterna forsteri'') is a tern in the family Laridae. The genus name ''Sterna'' is derived from Old English "stearn", "tern", and ''forsteri'' commemorates the naturalist Johann Reinhold Forster. It breeds inland in North Amer ...
or a
Mediterranean gull The Mediterranean gull (''Ichthyaetus melanocephalus'') is a small gull. The scientific name is from Ancient Greek. The genus ''Ichthyaetus'' is from ''ikhthus'', "fish", and ''aetos'', "eagle", and the specific ''melanocephalus'' is from ''m ...
. Juvenile gull-billed terns have a fainter mask, but otherwise look much like winter adults. Juvenile Sandwich terns have a short bill, and are frequently mistaken for gull-billed tern where the latter species is uncommon, such as
North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian S ...
coasts.


Distribution and habitat

It breeds in warmer parts of the world in southern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
, temperate and eastern
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
, both coasts of North America, eastern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
. This
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
has a number of geographical races, differing mainly in size and minor plumage details. All forms show a post-breeding dispersal, but the northern breeders are most
migratory Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
, wintering south to
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, the Caribbean and northern South America, southern Asia and New Zealand. The gull-billed tern is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' (
AEWA The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA) is an independent international treaty developed under the auspices of the United Nations Environment Programme's Convent ...
) applies.


Behaviour and ecology


Breeding

The gull-billed tern breeds in colonies on lakes, marshes and coasts (including bays and earthen levees). It nests in a ground scrape and lays two to five eggs. While widely distributed in freshwater areas in Eurasia, it is associated almost solely with saltwater, coastal areas in North America.


Food and feeding

This is a somewhat atypical tern, in appearance like a ''Sterna'' tern, but with feeding habits more like the ''Chlidonias'' marsh terns,
black tern The black tern (''Chlidonias niger'') is a small tern generally found in or near inland water in Europe, Western Asia and North America. As its name suggests, it has predominantly dark plumage. In some lights it can appear blue in the breeding s ...
and
white-winged tern The white-winged tern, or white-winged black tern (''Chlidonias leucopterus'' or ''Chlidonias leucoptera''), is a species of tern in the family Laridae. It is a small species generally found in or near bodies of fresh water across much of the wor ...
. It does not normally plunge dive for
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...
like the other white terns, and has a broader diet than most other terns. It largely feeds on insects taken in flight, and also often hunts over wet fields and even in brushy areas, to take amphibians and small mammals. It is also an opportunistic feeder, and has been observed to pick up and feed on dead dragonflies from the road.


Gallery

File:Tern in Chilka, Orissa I IMG 9309.jpg, Non-breeding in
Chilika Chilika Lake is a brackish water lagoon, spread over the Puri, khordha and Ganjam districts of Odisha state on the east coast of India, at the mouth of the Daya River, flowing into the Bay of Bengal, covering an area of over . It is the b ...
,
Odisha Odisha (English: , ), formerly Orissa ( the official name until 2011), is an Indian state located in Eastern India. It is the 8th largest state by area, and the 11th largest by population. The state has the third largest population of Sc ...
, India File:Tern in Chilka, Orissa I IMG 9354.jpg, Juvenile/ 1st winter in Chilika, Odisha, India File:Gelochelidon nilotica.jpg File:Gull-billed Tern (Gelochelidon nilotica) W IMG 6889.jpg, in
Krishna Wildlife Sanctuary Krishna Wildlife Sanctuary is a wildlife sanctuary and estuary located in Andhra Pradesh, India. It is one of the rarest eco-regions of the world because it harbors vast tracts of pristine mangrove forests. It is believed by conservationists to ...
,
Andhra Pradesh Andhra Pradesh (, abbr. AP) is a state in the south-eastern coastal region of India. It is the seventh-largest state by area covering an area of and tenth-most populous state with 49,386,799 inhabitants. It is bordered by Telangana to t ...
, India File:Gelochelidon nilotica vanrossemi.jpg, van Rossem's gull-billed terns at the San Diego National Wildlife Refuge Gull-billed terns (Gelochelidon nilotica).jpg,
Kazinga Channel The Kazinga Channel in Uganda is a wide, long natural channel that links Lake Edward and Lake George, and a dominant feature of Queen Elizabeth National Park. The channel attracts a varied range of animals and birds, with one of the world's l ...
,
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The south ...


References


External links

* * * * * * * * {{Taxonbar, from=Q18834
gull-billed tern The gull-billed tern (''Gelochelidon nilotica''), formerly ''Sterna nilotica'', is a tern in the family Laridae. It is widely distributed and breeds in scattered localities in Europe, Asia, northwest Africa, and the Americas. The Australian gull ...
Birds of the Americas Birds of the Dominican Republic Birds of Eurasia Birds described in 1789 Taxa named by Johann Friedrich Gmelin Articles containing video clips Cosmopolitan birds Holarctic birds