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''Stemonidium hypomelas'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
deep-sea The deep sea is broadly defined as the ocean depth where light begins to fade, at an approximate depth of or the point of transition from continental shelves to continental slopes. Conditions within the deep sea are a combination of low tempe ...
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 20 families, 164 genera, and about 1000 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
in the family
Serrivomeridae Sawtooth eels are a family, Serrivomeridae, of eels found in temperate and tropical seas worldwide. Sawtooth eels get their name from the human-like arrangement of inward-slanting teeth attached to the vomer bone in the roof of the mouth. They ...
, originally described from a specimen taken from a depth of around near Niihau Island,
Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
in 1902. It can be distinguished from the related sawtooth eels by its reduced
dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiology ...
, which more resemble those of the
snipe eel Snipe eels are a family, Nemichthyidae, of eels that consists of nine species in three genera. They are pelagic fishes, found in every ocean, mostly at depths of but sometimes as deep as . Depending on the species, adults may reach in length, y ...
. It is of no significance to
fisheries Fishery can mean either the enterprise of raising or harvesting fish and other aquatic life or, more commonly, the site where such enterprise takes place ( a.k.a., fishing grounds). Commercial fisheries include wild fisheries and fish farm ...
.


Distribution and habitat

''Stemonidium hypomelas'' is widely distributed in the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...
and the southern
Atlantic Ocean The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
, at depths of Scientific
trawl Trawling is an industrial method of fishing that involves pulling a fishing net through the water behind one or more boats. The net used for trawling is called a trawl. This principle requires netting bags which are towed through water to catch di ...
s conducted off
Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
show that this species occurs below during the day, with most between . At night, a small subset of the population migrates upward to a minimum depth of .


Description

''Stemonidium hypomelas'' has an elongated, band-shaped body, nearly of uniform depth in the middle and tapering towards the
head A head is the part of an organism which usually includes the ears, brain, forehead, cheeks, chin, eyes, nose, and mouth, each of which aid in various sensory functions such as sight, hearing, smell, and taste. Some very simple ani ...
and
tail The tail is the elongated section at the rear end of a bilaterian animal's body; in general, the term refers to a distinct, flexible appendage extending backwards from the midline of the torso. In vertebrate animals that evolution, evolved to los ...
. The dorsal profile from the back of the head to the base of the
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit ** podium * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ...
is nearly straight; the
jaw The jaws are a pair of opposable articulated structures at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth ...
s are long and taper to a very delicate, slender tip, with the
lower Lower may refer to: * ''Lower'' (album), 2025 album by Benjamin Booker *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is sit ...
slightly longer than the upper. The
teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
are reduced to small granular plates arranged in a
quincunx A quincunx ( ) is a geometry, geometric pattern consisting of five points arranged in a cross, with four of them forming a Square (geometry), square or rectangle and a fifth at its center. The same pattern has other names, including "in saltire" ...
on the jaws and
vomer The vomer (; ) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxillary bones. The vomer forms ...
with the pointed tips pointing backward. The
eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...
s are very small and located in the middle of the head. There are two pairs of slit-shaped
nostril A nostril (or naris , : nares ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbinates ...
s in front of the eyes, the anterior shorter than the posterior. The
gill A gill () is a respiration organ, respiratory organ that many aquatic ecosystem, aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow r ...
slits are long and very
oblique Oblique may refer to: * an alternative name for the character usually called a slash (punctuation) ( / ) *Oblique angle, in geometry * Oblique triangle, in geometry * Oblique lattice, in geometry * Oblique leaf base, a characteristic shape of the ...
. The long
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
and
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
s both begin about half a head length behind the gill opening. The first
fin ray Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only b ...
s of the dorsal fin are delicate and difficult to detect. The anal fin is somewhat taller than the dorsal, with the fin rays of both becoming longer and more crowded towards the tail. The tail is short and pointed, but not filamentous. There is no
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...
. The upper half of the body is light gray and covered with fine black specks of pigment, and the lower half is abruptly jet black. The head is entirely black, becoming lighter at the tips of the jaws and the back of the head, and the last sixth of the tail is also black. The fins are all lighter in color. The
male Male (Planet symbols, symbol: ♂) is the sex of an organism that produces the gamete (sex cell) known as sperm, which fuses with the larger female gamete, or Egg cell, ovum, in the process of fertilisation. A male organism cannot sexual repro ...
s are smaller than the
female An organism's sex is female ( symbol: ♀) if it produces the ovum (egg cell), the type of gamete (sex cell) that fuses with the male gamete (sperm cell) during sexual reproduction. A female has larger gametes than a male. Females and ...
s, with larger nasal rosettes and eyes, and sometimes flatter tails. The maximum known length is for a
male Male (Planet symbols, symbol: ♂) is the sex of an organism that produces the gamete (sex cell) known as sperm, which fuses with the larger female gamete, or Egg cell, ovum, in the process of fertilisation. A male organism cannot sexual repro ...
and for a
female An organism's sex is female ( symbol: ♀) if it produces the ovum (egg cell), the type of gamete (sex cell) that fuses with the male gamete (sperm cell) during sexual reproduction. A female has larger gametes than a male. Females and ...
.


Biology and ecology

Analysis of
stomach The stomach is a muscular, hollow organ in the upper gastrointestinal tract of Human, humans and many other animals, including several invertebrates. The Ancient Greek name for the stomach is ''gaster'' which is used as ''gastric'' in medical t ...
contents show that ''Stemonidium hypomelas'' feed on
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s, mostly
euphausiid Krill ''(Euphausiids)'' (: krill) are small and exclusively marine crustaceans of the order Euphausiacea, found in all of the world's oceans. The name "krill" comes from the Norwegian word ', meaning "small fry of fish", which is also often a ...
s but rarely also
shrimp A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...
s and
amphipod Amphipoda () is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods () range in size from and are mostly detritivores or scavengers. There are more than 10,700 amphipod species cur ...
s.
Reproduction Reproduction (or procreation or breeding) is the biological process by which new individual organisms – "offspring" – are produced from their "parent" or parents. There are two forms of reproduction: Asexual reproduction, asexual and Sexual ...
is
oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...
, with high reported fecundities ranging from 7,000 to almost 14,000
eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...
per female. Female length at maturation varies from , and the large size of their
gonad A gonad, sex gland, or reproductive gland is a Heterocrine gland, mixed gland and sex organ that produces the gametes and sex hormones of an organism. Female reproductive cells are egg cells, and male reproductive cells are sperm. The male gon ...
s relative to their bodies suggests that they may be
semelparous Semelparity and iteroparity are two contrasting reproductive strategies available to living organisms. A species is considered ''semelparous'' if it is characterized by a single reproduction, reproductive episode before death, and ''iteroparous ...
, investing a high proportion of their available resources into a single, large batch of eggs. The males mature at long; their better-developed nares and eyes may be an
adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the p ...
for finding widely scattered
mate Mate may refer to: Science * Mate, one of a pair of animals involved in: ** Mate choice, intersexual selection *** Mate choice in humans ** Mating * Multi-antimicrobial extrusion protein, or MATE, an efflux transporter family of proteins Pers ...
s in the open ocean.


References

{{Taxonbar, from=Q14743685, from2=Q2033783 Serrivomeridae Fish described in 1905 Taxa named by Charles Henry Gilbert Monotypic fish genera