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''Stellera'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
in the family
Thymelaeaceae The Thymelaeaceae are a cosmopolitan family of flowering plants composed of 50 genera (listed below) and 898 species.Zachary S. Rogers (2009 onwards)A World Checklist of Thymelaeaceae (version 1) Missouri Botanical Garden Website, St. Louis. It ...
, with a single species ''Stellera chamaejasme'' found in mountainous regions of
Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the former ...
, China,
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
and
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;; ...
. ''S. chamaejasme'' is a herbaceous
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
plant with heads of white, pink or yellow flowers, grown as an ornamental plant in
rock garden A rock garden, also known as a rockery and formerly as a rockwork, is a garden, or more often a part of a garden, with a landscaping framework of rocks, stones, and gravel, with planting appropriate to this setting. Usually these are small ...
s and
alpine house A greenhouse (also called a glasshouse, or, if with sufficient heating, a hothouse) is a structure with walls and roof made chiefly of transparent material, such as glass, in which plants requiring regulated climatic conditions are grown.These ...
s, but considered a
weed A weed is a plant considered undesirable in a particular situation, "a plant in the wrong place", or a plant growing where it is not wanted.Harlan, J. R., & deWet, J. M. (1965). Some thoughts about weeds. ''Economic botany'', ''19''(1), 16-24. ...
playing a rôle in the
desertification Desertification is a type of land degradation in drylands in which biological productivity is lost due to natural processes or induced by human activities whereby fertile areas become increasingly arid. It is the spread of arid areas caused b ...
of grasslands in parts of its native range. Like many others of its family, it is a
poisonous plant Plants that produce toxins are referred to as poisonous plants. Plants that cause irritation on contact are also described as "poisonous". The toxins in poisonous plants affect herbivores, and deter them from consuming the plants. Plants cannot ...
with medicinal and other useful properties.


Common names

Two common names recorded for the plant in
Mongolian Mongolian may refer to: * Something of, from, or related to Mongolia, a country in Asia * Mongolian people, or Mongols * Mongolia (1911–24), the government of Mongolia, 1911–1919 and 1921–1924 * Mongolian language * Mongolian alphabet * Mong ...
are одои далан туруу ''odoi dalan turuu'' and чонын Чолбодос ''choniin'' (="of the wolf") ''cholbodos'' (="?"), and a common name for the plant in
Tibetan Tibetan may mean: * of, from, or related to Tibet * Tibetan people, an ethnic group * Tibetan language: ** Classical Tibetan, the classical language used also as a contemporary written standard ** Standard Tibetan, the most widely used spoken diale ...
is ''rejag''.Medicinal Plants in Mongolia pub. World Health Organization, Regional Office for the Western Pacific Region 2013

/ref>


Description

''Stellera chamaejasme'' is a
herbaceous perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
. Unbranched stems, 20–30 cm tall, emerge in a cluster from an underground rhizome. Narrow, overlapping leaves are borne along the stems. Individual leaves are narrow and pointed, up to 2 cm long. The flowers are grouped into rounded tightly packed terminal heads. Flowers lack
petal Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usuall ...
s, instead having petaloid
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined ...
s forming a tube up to 1.5 cm long with usually five (but possibly four or six) short lobes. The flower colour varies from shades of pink and white to yellow. There are twice as many stamens as calyx lobes, in two series. The ovary has a single chamber (
locule A locule (plural locules) or loculus (plural loculi) (meaning "little place" in Latin) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usu ...
). The fruit is a dry
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part ( exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kerne ...
, enclosed by the remains of the calyx.


Taxonomy

The genus ''Stellera'' was first described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in 1753. He recognized two species: ''Stellera passerina'' (now placed in the genus ''
Thymelaea ''Thymelaea'' (English: the Sparrow-worts) is a genus of about 30 species of evergreen shrubs and herbs in the flowering plant family Thymelaeaceae, native to the Canary Islands, the Mediterranean region, north to central Europe, and east to ce ...
'' as '' T. passerina'') and ''Stellera chamaejasme''. The generic name ''Stellera'' (not to be confused with the entirely unrelated ''
Stellaria ''Stellaria'' is a genus of about 190 species of flowering plants in the family Caryophyllaceae, with a cosmopolitan distribution. Common names include starwort, stitchwort and chickweed. Description ''Stellaria'' species are relatively sm ...
'') commemorates
Georg Wilhelm Steller Georg Wilhelm Steller (10 March 1709 – 14 November 1746) was a German botanist, zoologist, physician and explorer, who worked in Russia and is considered a pioneer of Alaskan natural history.Evans, Howard Ensign. Edward Osborne Wilson (col.) ...
(Stöller), while the specific epithet ''chamaejasme'' is a rendering into botanical Latin orthography of the Greek χαμαί ''khamai'' "(down) on the ground" and ιασμε ''iasme'' "
jasmine Jasmine ( taxonomic name: ''Jasminum''; , ) is a genus of shrubs and vines in the olive family ( Oleaceae). It contains around 200 species native to tropical and warm temperate regions of Eurasia, Africa, and Oceania. Jasmines are widely cult ...
". The name in its entirety thus means "Steller's plant that resembles a kind of jasmine (that creeps) on the ground". The flower of ''Stellera chamaejasme'' is fragrant like that of jasmine and also has a wine-red exterior, like that of certain species of jasmine, e.g. common jasmine ''
Jasminum officinale ''Jasminum officinale'', known as the common jasmine or simply jasmine, is a species of flowering plant in the olive family Oleaceae. It is native to the Caucasus and parts of Asia, also widely naturalized. It is also known as summer jasmine, ...
'' and the Chinese species '' Jasminum polyanthum''. Unlike jasmine, however, ''Stellera'' is a herbaceous plant, not a woody one, and its stems do not twine. Many species names were later created in the genus, but all are now usually considered synonyms of other species, including ''S. chamaejasme'', although the ''Flora of China'' states that there are 10 to 12 species. Studies in 2002 and 2009, based on
chloroplast DNA Chloroplast DNA (cpDNA) is the DNA located in chloroplasts, which are photosynthetic organelles located within the cells of some eukaryotic organisms. Chloroplasts, like other types of plastid, contain a genome separate from that in the cell n ...
, placed ''Stellera'' in a small group of related genera, either as sister to ''
Wikstroemia ''Wikstroemia'' is a genus of 55-70 species of flowering shrubs and small trees in the mezereon family, Thymelaeaceae. Hawaiian species are known by the common name ‘ākia. Medicinal uses ''Wikstroemia indica'' () is one of the 50 fundamen ...
'' or embedded within it; however for most genera only one species was included.


Distribution and habitat

''Stellera chamaejasme'' is native to northern and western China, Tibet, the Himalayas (Nepal, Bhutan), the state of
Uttar Pradesh Uttar Pradesh (; , 'Northern Province') is a state in northern India. With over 200 million inhabitants, it is the most populated state in India as well as the most populous country subdivision in the world. It was established in 1950 ...
in north India, Russia and Mongolia. In China, it is found on sunny dry slopes and sandy places between 2600 and 4200 m.


Cultivation

''Stellera chamaejasme'' is cultivated as an ornamental plant in
rock garden A rock garden, also known as a rockery and formerly as a rockwork, is a garden, or more often a part of a garden, with a landscaping framework of rocks, stones, and gravel, with planting appropriate to this setting. Usually these are small ...
s and
alpine house A greenhouse (also called a glasshouse, or, if with sufficient heating, a hothouse) is a structure with walls and roof made chiefly of transparent material, such as glass, in which plants requiring regulated climatic conditions are grown.These ...
s. It is considered difficult to grow, needing a sunny position and gritty soil if grown outside, or a large pot if grown under cover. It is propagated by seed.


Toxicity

The
Russians , native_name_lang = ru , image = , caption = , population = , popplace = 118 million Russians in the Russian Federation (2002 ''Winkler Prins'' estimate) , region1 = , pop1 ...
living in
Dauria Transbaikal, Trans-Baikal, Transbaikalia ( rus, Забайка́лье, r=Zabaykalye, p=zəbɐjˈkalʲjɪ), or Dauria (, ''Dauriya'') is a mountainous region to the east of or "beyond" (trans-) Lake Baikal in Far Eastern Russia. The steppe and ...
still hold Stellera in high esteem on account of its root, despite the fact that its violent effects have already dispatched a good many people to the afterlife. This root resembles a crudely-carved human figure to a degree even greater than that of the
famous Famous may refer to: Companies * Famous Brands, a South African restaurant franchisor * Famous Footwear, an American retail store chain * Famous Music, the music publishing division of Paramount Pictures * Famous Studios, the animation division ...
''Alraune'' or Mandragora, such that one can often distinguish clearly in its natural form rotuberances resemblinga head, arms and legs; which has led to its being given the most apposite name of ''Muzhik koren'' �ужик коренor "Man root" by the Russians. The oldest rootstocks of this plant can reach the size of a large carrot, and f taken as medicineproduce the most violent effects. Such old roots can produce more than fifty - and sometimes as many as a hundred - flowering shoots, which, crowned with their beautiful and fragrant flowers, give not the least hint of the violent and pernicious effects residing in the root that bore them. The exterior of the flower is usually of a dark reddish-purple, or, more rarely
sulphur Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...
-yellow colour; while the interior is white. Add to this the fact that the flowers display, upon opening, a white border and in the middle a red or yellow patch, and the flowers present the most
ravishing ''Ravishing'' (French: ''Ravissante'') is a 1960 French- Italian comedy film directed by Robert Lamoureux and starring Lamoureux, Sylva Koscina and Philippe Noiret.Oscherwitz & Higgins p.143 It was shot at the Epinay Studios in Paris. The fi ...
aspect. The young
Tungus Tungusic peoples are an ethno-linguistic group formed by the speakers of Tungusic languages (or Manchu–Tungus languages). They are native to Siberia and Northeast Asia. The Tungusic phylum is divided into two main branches, northern (Evenic or ...
boys are in the habit of adorning their bare heads with a kind of hat which they create most artistically by interweaving whole flowering stems of the plant. On this plant one often finds a flower in which two normal flowers seem fused into one, bearing a corolla with nine lobes and eighteen stamens arranged in two rows.

ranslated from the French of a text closely based on the account of Prussian naturalist and explorer Peter Simon Pallas ].
The plant is virulently poisonous and has caused fatalities both in humans and in livestock. The powdered roots have been used as a laxative, as a pesticide and as a Fish toxins, fish poison, and have also been used in small doses as a drastic
anthelmintic Anthelmintics or antihelminthics are a group of antiparasitic, antiparasitic drugs that expel parasitic worms (helminths) and other internal parasites from the body by either stunning or killing them and without causing significant damage to the h ...
for sheep and goats. The plant is common in Western China, where it goes by the common name of ''Langdu'' (狼毒花) lit. "wolf poison" (狼 ''lang'' "wolf" + 毒 ''dú'' "poison" + 花 ''huā'' "flower"). It is used as a medicinal herb in China, but can be considered an undesirable element in the flora if it should proliferate to too great an extent, as its large, water-thirsty roots speed up the desertification of prairies. A work on native Chinese medicinal plants aimed at farmers states that ''Stellera'' is a very poisonous plant used as an insecticide and that, if consumed by an animal, will cause the victim's
intestines The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans ...
to disintegrate. Corroboration of this evidence for the damaging effect upon the intestines of certain plants belonging to the Thymelaceae may be found in an account of "
Lasiosiphon kraussianus ''Lasiosiphon'' is a genus of flowering plants belonging to the family Thymelaeaceae. Its native range is tropical and southern Africa, Madagascar, southwestern Arabian Peninsula, India, Sri Lanka. Species: *'' Lasiosiphon ambondrombensis'' ...
Hutch. & Dalz. " (referable, possibly to ''Lasiosiphon kraussianus'' (Meisn.) Meisn. or a ''Gnidia'' sp.) of
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
: the plant is exceedingly poisonous and rapidly fatal to stock: the intestines of an animal perforate about a day after eating it. This lethal property is put to use by certain African tribes who use the powdered root of the plant to poison waterholes during the rainy season, the poison remaining potent for seven days and killing any animal which drinks it. The plant family to which ''Stellera'' belongs - Thymelaceae - is notable for the number of poisonous species which it contains and also for a certain similarity in chemistry to the family
Euphorbiaceae Euphorbiaceae, the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of a genus in the family. Most spurges, such as '' Euphorbia paralias'', are herbs, but some, ...
, both families having a number of genera producing
phorbol ester Phorbol esters are a class of chemical compounds found in a variety of plants, particularly in the families Euphorbiaceae and Thymelaeaceae. Chemically, they are ester derivatives of the tetracyclic diterpenoid phorbol. Biological activity Protein ...
s. It is interesting to note, in this context, that Chinese herbal medicine recognises a similarity in action between ''Stellera'' and certain ''
Euphorbia ''Euphorbia'' is a very large and diverse genus of flowering plants, commonly called spurge, in the family Euphorbiaceae. "Euphorbia" is sometimes used in ordinary English to collectively refer to all members of Euphorbiaceae (in deference to t ...
'' species: Perry (1980) notes that, in a Chinese materia medica of 1959, ''Stellera'', ''
Euphorbia fischeriana ''Euphorbia'' is a very large and diverse genus of flowering plants, commonly called spurge, in the family Euphorbiaceae. "Euphorbia" is sometimes used in ordinary English to collectively refer to all members of Euphorbiaceae (in deference to t ...
''
Steud. Ernst Gottlieb von Steudel (30 May 1783 – 12 May 1856) was a German physician and an authority on grasses. Biography Ernst Gottlieb von Steudel was born at Esslingen am Neckar in Baden-Württemberg. He was educated at the University of Tübinge ...
(syn. E. pallasii
Turcz. Nikolai Stepanovich Turczaninow ( ru , Николай Степанович Турчанинов, 1796 in Nikitovka, now in Krasnogvardeysky District, Belgorod Oblast, Russia – 1863 in Kharkov) was a Russian botanist and plant collector wh ...
) and ''
Euphorbia sieboldiana ''Euphorbia sieboldiana'', the Siebold's spurge, is a species of flowering plant in the spurge family Euphorbiaceae. It is native to eastern Asia, where it is found in China, Japan, Korea, and eastern Russia.Morr. &
Decne. Joseph Decaisne (7 March 1807 – 8 January 1882) was a French botanist and agronomist. He became an ''aide-naturaliste'' to Adrien-Henri de Jussieu (1797-1853), who served as the chair of rural botany. It was during this time that he began to st ...
are listed under the same heading (no. 86, ''langdu'') - and as possessing the same or very similar medicinal properties: pungent, poisonous plants used as
cathartic In medicine, a cathartic is a substance that ''accelerates'' defecation. This is similar to a laxative, which is a substance that ''eases'' defecation, usually by softening feces. It is possible for a substance to be both a laxative and a cathart ...
s, anthelmintics,
expectorant Mucoactive agents are a class of chemical agents that aid in the clearance of mucus or sputum from the upper and lower airways, including the lungs, bronchi, and trachea. Mucoactive drugs include expectorants, mucolytics, mucoregulators, and mucok ...
s, also used topically to treat ulcers and
skin diseases A skin condition, also known as cutaneous condition, is any medical condition that affects the integumentary system—the organ system that encloses the body and includes skin, nails, and related muscle and glands. The major function of this s ...
.


Chemistry and properties

The principal constituents of ''Stellera chamaejasme'' include, among others,
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s,
coumarin Coumarin () or 2''H''-chromen-2-one is an aromatic organic chemical compound with formula . Its molecule can be described as a benzene molecule with two adjacent hydrogen atoms replaced by a lactone-like chain , forming a second six-membered ...
s,
lignan The lignans are a large group of low molecular weight polyphenols found in plants, particularly seeds, whole grains, and vegetables. The name derives from the Latin word for "wood". Lignans are precursors to phytoestrogens. They may play a rol ...
s and
diterpenoid Diterpenes are a class of chemical compounds composed of four isoprene units, often with the molecular formula C20H32. They are biosynthesized by plants, animals and fungi via the HMG-CoA reductase pathway, with geranylgeranyl pyrophosphate being ...
s. A recent work on the medicinal plants of Mongolia notes the presence in the root (rhizome) of
sugars Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose, fructose, and galactose. Compound sugars, also called disaccharides or double ...
,
organic acids An organic acid is an organic compound with acidic properties. The most common organic acids are the carboxylic acids, whose acidity is associated with their carboxyl group –COOH. Sulfonic acids, containing the group –SO2OH, are ...
,
saponin Saponins (Latin "sapon", soap + "-in", one of), also selectively referred to as triterpene glycosides, are bitter-tasting usually toxic plant-derived organic chemicals that have a foamy quality when agitated in water. They are widely distributed ...
s and
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner ...
s and the following specific compounds: the flavonoids 5,7-dihydroxy-4',11-dimethoxy-3',14-dimethylbenzoflavanone, ruixianglangdusu A and B, 4',4'",5,5",7,7"-hexahydroxy-3,3"-biflavone, 7-methoxyneochamaejasmin A; the coumarins: sfondine, isobergapten, pimpinellin, isopimpinellin,
umbelliferone Umbelliferone, also known as 7-hydroxycoumarin, hydrangine, skimmetine, and ''beta''-umbelliferone, is a natural product of the coumarin family. It absorbs ultraviolet light strongly at several wavelengths. There are some indications that this ...
, daphniretin, bicoumastechamin and daphnetin; diterpenes (unspecified); the lignans: (+)-kusunokinin, lirioresinol-B, magnolenin C, (-)-pinoresinol monomethyl ether, (-)-
pinoresinol Pinoresinol is a tetrahydrofuran lignan found in ''Styrax sp.'', '' Forsythia suspensa, and in Forsythia koreana''. It is also found in the caterpillar of the cabbage butterfly, ''Pieris rapae'' where it serves as a defence against ants. In food, ...
, (+)-
matairesinol Matairesinol is an organic compound. It is classified as a lignan, i.e., a type of phenylpropanoid. It is present in some cereals, e.g. rye, and together with Secoisolariciresinol, has attracted much attention for its beneficial nutritional effec ...
, isohinokinin, and (-)-eudesmin; and the steroids:
daucosterol Daucosterol ( eleutheroside A) is a natural phytosterol-like compound. It is the glucoside of β-sitosterol β-sitosterol (beta-sitosterol) is one of several phytosterols (plant sterols) with chemical structures similar to that of cholestero ...
,
β-sitosterol β-sitosterol (beta-sitosterol) is one of several phytosterols (plant sterols) with chemical structures similar to that of cholesterol. It is a white, waxy powder with a characteristic odor, and is one of the components of the food additive E49 ...
. Above-ground parts of the plant were found to contain the coumarins: daphnorin, daphnetin,
daphnoretin Daphnoretin is a protein kinase C activator isolated from '' Wikstroemia indica'' C.A. Mey, one of the 50 fundamental herbs in traditional Chinese medicine. It has also been found to occur in ''Stellera chamaejasme'' - the single species in a genu ...
, daphnetin 8-O-b-D-glycopyranoside and chamaejasmoside. A scientific paper of 2015 refers to this plant - regarded as a choice and hard-to-grow ornamental by European and American gardeners - as being one of the most toxic of grassland
weed A weed is a plant considered undesirable in a particular situation, "a plant in the wrong place", or a plant growing where it is not wanted.Harlan, J. R., & deWet, J. M. (1965). Some thoughts about weeds. ''Economic botany'', ''19''(1), 16-24. ...
s in the range where it is native and notes that cattle which consume its shoots and flowers may be fatally poisoned. The paper notes further that populations of the plant are in no way endangered, having been flourishing and increasing for some years: this appears to be due not simply to the plant's competing vigorously with other species for water and nutrients, but also to its containing / secreting herbicidal compounds.Water and
ethanol Ethanol (abbr. EtOH; also called ethyl alcohol, grain alcohol, drinking alcohol, or simply alcohol) is an organic compound. It is an alcohol with the chemical formula . Its formula can be also written as or (an ethyl group linked to a h ...
extracts of S. chamaejasme inhibited seed germination and/or seedling growth in no fewer than 13 plant species, and the phytotoxic effects were stronger upon dicotyledonous plants than upon
monocotyledon Monocotyledons (), commonly referred to as monocots, ( Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of ...
ous plants. The phytotoxic compounds were observed to be liberated particularly by dead or moribund specimens of ''S. chamaejasme'' and to lead to reduced seedling growth in the
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in law ...
es ''
Lolium perenne ''Lolium perenne'', common name perennial ryegrass, English ryegrass, winter ryegrass, or ray grass, is a grass from the family Poaceae. It is native to Europe, Asia and northern Africa, but is widely cultivated and naturalised around the world. ...
'' L., ''
Psathyrostachys juncea ''Psathyrostachys juncea'' is a species of grass known by the common name Russian wildrye. It was formerly classified as ''Elymus junceus''. It is native to Russia and China, and has been introduced to other parts of the world, such as Canada an ...
'' (Fisch.) Nevski and ''
Bromus inermis ''Bromus inermis'' is a species of the true grass family ( Poaceae). This rhizomatous grass is native to Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and th ...
'' Leyss. and the
legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock fo ...
s ''
Melilotus suaveolens ''Melilotus'', known as melilot, sweet clover, and kumoniga (from the Cumans),Bulgarian Folk Customs, Mercia MacDermott, pg 27 is a genus in the family Fabaceae (the same family that also includes the ''Trifolium'' clovers). Members are known ...
'' Ledeb. (see ''
Melilotus ''Melilotus'', known as melilot, sweet clover, and kumoniga (from the Cumans),Bulgarian Folk Customs, Mercia MacDermott, pg 27 is a genus in the family Fabaceae (the same family that also includes the ''Trifolium'' clovers). Members are known ...
''), ''
Onobrychis viciifolia ''Onobrychis viciifolia'', also known as ''O. sativa'' or common sainfoin () has been an important forage legume in temperate regions until the 1950s. During the Green Revolution it was replaced by high yielding alfalfa and clover species. Due ...
'' Scop. (sainfoin) and ''
Medicago sativa Alfalfa () (''Medicago sativa''), also called lucerne, is a perennial flowering plant in the legume family Fabaceae. It is cultivated as an important forage crop in many countries around the world. It is used for grazing, hay, and silage, as ...
'' L. (alfalfa). Furthermore, pesticidal properties were confirmed to be present in ''S. chameajasme'': the ethanolic extract of S. chameajasme strongly inhibited the growth of the following insect pests: the butterfly ''
Pieris rapae ''Pieris rapae'' is a small- to medium-sized butterfly species of the whites-and-yellows family Pieridae. It is known in Europe as the small white, in North America as the cabbage white or cabbage butterfly, on several continents as the small c ...
'', the aphid ''
Myzus persicae ''Myzus persicae'', known as the green peach aphid, greenfly, or the peach-potato aphid, is a small green aphid belonging to the order Hemiptera. It is the most significant aphid pest of peach trees, causing decreased growth, shrivelling of the ...
'' and the corn-borer moth ''
Ostrinia furnacalis ''Ostrinia furnacalis'' is a species of moth in the family Crambidae, the grass moths. It was described by Achille Guenée in 1854 and is known by the common name Asian corn borer since this species is found in Asia and feeds mainly on corn crop ...
'', and showed contact and oral toxicities against two other stem-borer moths which are pests of
rice Rice is the seed of the grass species '' Oryza sativa'' (Asian rice) or less commonly '' Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera '' Zizania'' and ''Porteresia'', both wild and domestica ...
: ''
Sesamia inferens ''Sesamia inferens'', the Asiatic pink stem borer, gramineous stem borer, pink borer, pink rice borer, pink rice stem borer, pink stem borer, purple borer, purple stem borer or purplish stem borer, is a moth of the family Noctuidae. The species ...
'' and ''
Chilo suppressalis ''Chilo suppressalis'', the Asiatic rice borer or striped rice stemborer, is a moth of the family Crambidae. It is a widespread species, known from Iran, India, Sri Lanka, China, eastern Asia, Japan, Taiwan, Malaysia to the Pacific. It is a se ...
''.


Papermaking

In
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
, the thick, fibrous root of ''Stellera'' is harvested, cooked, and beaten for making paper.


References

{{Taxonbar, from=Q3487244 Thymelaeoideae Monotypic Malvales genera Medicinal plants of Asia Garden plants of Asia Poisonous plants