The steenbok (''Raphicerus campestris'') is a common small
antelope
The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia.
Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...
of southern and eastern
Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. It is sometimes known as the steinbuck or steinbok.
Description
Steenbok resemble small
oribi, standing 45–60 cm (16"–24") at the shoulder, and average ~12 kg. Their
coat
A coat typically is an outer garment for the upper body as worn by either gender for warmth or fashion. Coats typically have long sleeves and are open down the front and closing by means of buttons, zippers, hook-and-loop fasteners, toggles, a ...
is any shade from fawn to rufous, typically rather orange. The underside, including chin and throat, is white, as is the ring around the eye. Ears are large with "finger-marks" on the inside. Males have straight, smooth, parallel
horns Horns or The Horns may refer to:
* Plural of Horn (instrument), a group of musical instruments all with a horn-shaped bells
* The Horns (Colorado), a summit on Cheyenne Mountain
* ''Horns'' (novel), a dark fantasy novel written in 2010 by Joe Hill ...
7–19 cm long (see image left). There is a black crescent-shape between the ears, a long black bridge to the glossy black nose, and a black circular scent-gland in front of the eye. The tail is not usually visible, being only 4–6 cm long.
Distribution
There are two distinct clusters in steenbok distribution. In
East Africa
East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa:
Due to the historical ...
, it occurs in central and southern
Kenya
)
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, ...
and northern
Tanzania
Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
. It was formerly widespread in
Uganda
}), is a landlocked country in East Africa
East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
,
[ but is now almost certainly extinct there. In ]Southern Africa
Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
, it occurs in Angola
, national_anthem = " Angola Avante"()
, image_map =
, map_caption =
, capital = Luanda
, religion =
, religion_year = 2020
, religion_ref =
, coordina ...
, Namibia
Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
, South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, Eswatini
Eswatini ( ; ss, eSwatini ), officially the Kingdom of Eswatini and formerly named Swaziland ( ; officially renamed in 2018), is a landlocked country in Southern Africa. It is bordered by Mozambique to its northeast and South Africa to its no ...
, Botswana
Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label=Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalahar ...
, Mozambique
Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...
, Zambia
Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most cent ...
, Zimbabwe
Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and Mozam ...
and probably Lesotho
Lesotho ( ), officially the Kingdom of Lesotho, is a country landlocked country, landlocked as an Enclave and exclave, enclave in South Africa. It is situated in the Maloti Mountains and contains the Thabana Ntlenyana, highest mountains in Sou ...
.
Habitat
Steenbok live in a variety of habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s from semi-desert
A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
, such as the edge of the Kalahari Desert and Etosha National Park
Etosha National Park is a national park in northwestern Namibia and one of the largest national parks in Africa. It was proclaimed a game reserve in March 1907 in Ordinance 88 by the Governor of German South West Africa, Friedrich von Lindequist. ...
, to open woodland and thickets, including open plain
In geography, a plain is a flat expanse of land that generally does not change much in elevation, and is primarily treeless. Plains occur as lowlands along valleys or at the base of mountains, as coastal plains, and as plateaus or uplands ...
s, stony savanna
A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
h, and ''Acacia
''Acacia'', commonly known as the wattles or acacias, is a large genus of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa and Australasia. The genus na ...
''–grassland
A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
mosaic
A mosaic is a pattern or image made of small regular or irregular pieces of colored stone, glass or ceramic, held in place by plaster/mortar, and covering a surface. Mosaics are often used as floor and wall decoration, and were particularly pop ...
s. They are said to favour unstable or transitional habitats.[Kingdon, Jonathan. 1997. ''The Kingdon Field Guide to African Mammals''. Academic Press, San Diego & London. Pp. 387–388. ()] At least in the central part of Kruger National Park
Kruger National Park is a South African National Park and one of the largest game reserves in Africa. It covers an area of in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends from north to south and from ea ...
, South Africa, Steenbok show a distinct preference for '' Acacia tortilis'' savannah throughout the year, with no tendency to migrate to moister areas during the dry season (unlike many larger African savannah ungulates, including species sympatric
In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species sh ...
with Steenbok in the wet season).
Population density is typically 0.3–1.0 individuals per square kilometre, reaching 4 per km2 in optimal habitats.[Kleiman, David G. ''et al.'', Eds. 2003. ''Grzimek's Animal Life Encyclopedia'', 2nd edn. Vol. 16: Mammals V. Gale Cengage Learning. Pp. 59–72.]
Diet
Steenbok typically browse
Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
on low-level vegetation (they cannot reach above 0.9 m), but are also adept at scraping up roots and tubers. In central Kruger National Park, Steenbok show a distinct preference for forbs, and then woody plants (especially '' Flueggea virosa'') when few forbs are available.[ They will also take fruits and only very rarely graze on grass.][ They are almost entirely independent of drinking water, gaining the moisture they need from their food.
]
Behaviour
Steenbok are active during the day and the night; however, during hotter periods, they rest under shade during the heat of the day. The time spent feeding at night increases in the dry season. While resting, they may be busy grooming, ruminating or taking brief spells of sleep.[
]
Anti-predator
At the first sign of trouble, steenbok typically lie low in the vegetation. If a predator or perceived threat comes closer, a steenbok will leap away and follow a zigzag route to try to shake off the pursuer. Escaping steenbok frequently stop to look back, and flight is alternated with prostration during extended pursuit. They are known to take refuge in the burrows of aardvarks
The aardvark ( ; ''Orycteropus afer'') is a medium-sized, burrowing, nocturnal mammal native to Africa. It is the only living species of the order Tubulidentata, although other prehistoric species and genera of Tubulidentata are known. Unlike ...
. Known predators include Southern African wildcat
The Southern African wildcat (''Felis lybica cafra'') is an African wildcat subspecies native to Southern and Eastern Africa.
In 2007, it was tentatively recognised as a distinct subspecies on the basis of genetic analysis. Morphological evidenc ...
, caracal
The caracal (''Caracal caracal'') () is a medium-sized wild cat native to Africa, the Middle East, Central Asia, and arid areas of Pakistan and northwestern India. It is characterised by a robust build, long legs, a short face, long tufted e ...
, jackals
Jackals are medium-sized canids native to Africa and Eurasia. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed ...
, leopard
The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
, martial eagle
The martial eagle (''Polemaetus bellicosus'') is a large eagle native to sub-Saharan Africa.Ferguson-Lees & Christie, ''Raptors of the World''. Houghton Mifflin Company (2001), . It is the only member of the genus ''Polemaetus''. A species of t ...
and pythons.
Breeding
Steenbok are typically solitary, except for when a pair come together to mate
Mate may refer to:
Science
* Mate, one of a pair of animals involved in:
** Mate choice, intersexual selection
** Mating
* Multi-antimicrobial extrusion protein, or MATE, an efflux transporter family of proteins
Person or title
* Friendship ...
. However, it has been suggested[ that pairs occupy consistent ]territories
A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal.
In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
while living independently, staying in contact through scent marking
In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...
s, so that they know where their mate is most of the time. Scent marking is primarily through dung midden
Dung middens, also known as ''dung hills'',''The New Encyclopaedia of Mammals'' D MacDonald 2002 Oxford are piles of dung that mammals periodically return to and build up. They are used as a form of territorial marker. A range of animals are know ...
s. Territories range from 4 hectares to 1 square kilometre. The male is aggressive during the female's oestrus
The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous ...
, engaging in "bluff-and-bluster" type displays with rival males—prolonged contests invariably involve well-matched individuals, usually in their prime.[Cohen, Michael. 1976. The Steenbok: A neglected species. ''Custos'' (April 1976): 23–26.]
Breeding occurs throughout the year, although more fawns are born November to December in the southern spring–summer; some females may breed twice a year. Gestation period
In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once it ...
is about 170 days, and usually a single precocial fawn is produced. The fawn is kept hidden in vegetation for 2 weeks, but suckles for 3 months. Females become sexually mature at 6–8 months and males at 9 months.
Steenbok are known to live for 7 years or more.
Taxonomy
Two subspecies
In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized: ''R. c. campestris'' in Southern Africa and ''R. c. neumanni'' of East Africa; although MSW3
''Mammal Species of the World: A Taxonomic and Geographic Reference'' is a standard reference work in mammalogy giving descriptions and bibliographic data for the known species of mammals. It is now in its third edition, published in late 2005, ...
also recognizes ''capricornis'' and ''kelleni''. Up to 24 subspecies have been described from Southern Africa, distinguished on such features as coat colour.
Gallery
Steenbok female.jpg , Female in South Africa
Steenbok (Raphicerus campestris) male.jpg, Backlit male showing white fur in ears, Tswalu Kalahari Res.
Steenbok (Raphicerus campestris) running composite.jpg, Female running in Damaraland
Damaraland was a name given to the north-central part of what later became Namibia, inhabited by the Damara (people), Damaras. It was bounded roughly by Ovamboland in the north, the Namib Desert in the west, the Kalahari Desert in the east, a ...
, Namibia
Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
Notes
References
External links
Wild About You (info and image)
{{Taxonbar, from=Q368033
Dwarf antelopes
Fauna of East Africa
Mammals of Angola
Mammals of Botswana
Mammals of Kenya
Mammals of Mozambique
Mammals of Namibia
Mammals of South Africa
Mammals of Eswatini
Mammals of Tanzania
Mammals of Zambia
Mammals described in 1811