Staphylininae are a

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

rove beetle The rove beetles are a family (biology), family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With over 66,000 species in thousand ...

s (

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Staphylinidae). They contain the typical rove beetles with their long but fairly robust blunt-headed and -tipped bodies and short

elytra An elytron (; ; : elytra, ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometime ...

, as well as some more unusually-shaped lineages.

Systematics

As it seems, the Staphylininae are part of a large

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

together with the subfamilies Euaesthetinae, Leptotyphlinae,

Megalopsidiinae ''Megalopinus'' is the only genus in the subfamily Megalopsidiinae of the Rove beetle, Staphylinidae. Species in this genus have large eyes, antennae with distinct di- or trisegmented clubs. The tarsal formula is 5-5-5. They have unique elongated ...

, Oxyporinae,

Paederinae The Paederinae are a subfamily of the Staphylinidae, rove beetles. , three tribes are accepted within this subfamily: Lathrobiini, Paederini, and Pinophilini. This insect is commonly known as Tomcat. Three of the genera of a subtribe of the ...

, Pseudopsinae, Scydmaeninae, Solieriinae, Steninae, and the

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

Protactinae which are only known from

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s. The numerous Staphylininae

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

are divided into six

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...

s, listed below along with some notable genera. However, a scientific study in 2020 proposed moving all tribes except Staphylinini to other subfamilies and raising several of Staphylinini's subtribes to tribal level. This would result in a new total of twelve tribes: Acylophorini, Afroquediini, Amblyopinini, Antimerini, †Baltognathini, Cyrtoquediini, Erichsoniini, Hyptiomini, Indoquediini, Quediini, Staphylinini and Tanygnathinini. * Coomaniini Żyła & Solodovnikov, 2019 ** '' Coomania'' Cameron, 1939 * Diochini Casey, 1906 ** '' Antarctothius'' Coiffait & Saiz ** '' Diochus'' Erichson, 1839 *** ''

Diochus electrus ''Diochus electrus'' is an extinct species of rove beetle in genus ''Diochus'', the only definitive fossil species in subfamily Staphylininae. The species is known only from the middle Eocene, Lutetian, Lutetian stage Baltic amber found in the B ...

'' Chatzimanolis & Engel, 2011 (

Lutetian The Lutetian is, in the geologic timescale, a stage (stratigraphy), stage or age (geology), age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it ...

Baltic amber Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the re ...

) * Maorothiini Diochus electrus ZooKeys-138-065-g001a

* Othiini C.G.Thomson, 1859 * Platyprosopini Lynch, 1884 * Staphylinini Latreille, 1802 (including Philonthini, Tanygnathinini) ** '' Cafius'' ** ''

Creophilus ''Creophilus'' is a genus of beetles of the Staphylinidae family (biology), family, subfamily Staphylininae. It includes some of the largest rove beetle species, up to 30 mm long. Almost all species live primarily on carrion and feed on maggots, ...

'' *** '' Creophilus maxillosus'' – hairy rove beetle ** '' Eucibdelus'' ** '' Gabrius'' ** '' Jurecekia'' ** '' Ocypus'' *** '' Ocypus olens'' – devil's coach-horse beetle *** '' Ocypus ophthalmicus'' ** '' Ontholestes'' ** '' Paederomimus'' ** '' Philonthus'' *** '' Philonthus cognatus'' ** '' Quedius'' *** '' Quedius curtipennis'' *** '' Quedius fuliginosus'' ** '' Rhynchocheilus'' ** '' Staphylinus'' ** '' Stevensia'' ** '' Thinopinus'' –

pictured rove beetle The pictured rove beetle (''Thinopinus pictus'') is a wingless rove beetle which lives on the sandy beaches of the West Coast of the United States from southern Alaska to Baja California. It is nocturnal, emerging at night from temporary sand bur ...

** ''

Velleius Marcus Velleius Paterculus (; ) was a Roman historian, soldier and senator. His Roman history, written in a highly rhetorical style, covered the period from the end of the Trojan War to AD 30, but is most useful for the period from the death o ...

'' Leach, 1819 *** '' Velleius dilatatus'' (Fabricius, 1787) * Xantholinini Erichson, 1839 ** '' Gyrohypnus'' Leach, 1819 ** '' Leptacinus'' Erichson, 1839 *** '' Leptacinus intermedius'' Donisthorpe, 1935 ** '' Xantholinus'' Erichson, 1839

References

External links

* * {{Staphylinidae-stub