''Squalius pyrenaicus'' , the Southern Iberian chub or Tagus chub, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of freshwater

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Leuciscidae Leuciscidae is a family of freshwater ray-finned fishes, formerly classified as a subfamily of the Cyprinidae, which contains the true minnows. Members of the Old World (OW) clade of minnows within this subfamily are known as European minnow ...

, which includes the daces, Eurasian minnows and related fishes. This species is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...

Taxonomy

''Squalius pyrenaicus'' was first formally described as ''Leuciscus pyrenaicus'' in 1868 by the German-born British

herpetologist Herpetology (from Ancient Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is a branch of zoology concerned with the study of amphibians (including frogs, salamanders, and caecilians (Gymnophiona)) and reptiles (in ...

and

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...

Albert Günther Albert Karl Ludwig Gotthilf Günther , also Albert Charles Lewis Gotthilf Günther (3October 18301February 1914), was a German-born British zoologist, ichthyologist, and herpetologist. Günther is ranked the second-most productive reptile tax ...

with its type locality given as the Mondego and Cintra rivers in Portugal. The Tagus chub belongs to the genus ''

Squalius ''Squalius'' is a genus of freshwater ray-finned fish belonging to the family (biology), family Leuciscidae, the daces, Eurasian minnows and reklated fishes. The fishies in this genus are found in Europe and Asia. Hybrid (biology), Hybridization ...

'', commonly referred to as chubs, which belongs to the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Leuciscinae Leuciscinae is a subfamily of freshwater Actinopterygii, ray-finned fishes belonging to the Family (biology), family Leuciscidae, which includes the fishes known as daces, chubs, shiners and minnows. The fishes in this subfamily are mainly found i ...

of the family Leuciscidae. This species is one of the parent species of two species of Iberian endemic chub '' S. alburnoides'' and the extinct '' S. palaciosi'', the other parent taxon of these species is unknown and may be extinct. Populations of chub in the

Sado River The Sado () is a river in southern Portugal; it is one of the major rivers in the country. It flows in a northerly direction (the only major Portuguese river to do so) through from its springs in the hills of Ourique before entering the Atlanti ...

catchment have been thought to be this species but have recently been identified as being closer to '' S. tartessicus.

Etymology

''Squalius pyrenaicus'' belongs to the genus ''Squalius'', this name was proposed by the French bioogist Charles Lucien Bonaparte in 1837 for a subgenus of the genus ''

Leuciscus ''Leuciscus'' is a genus of freshwater and brackish water ray-finned fishes belonging to the family Leuciscidae, which includes the daces, Eurasian minnows and related species. They are inland water fishes commonly called Eurasian daces. Th ...

'' for the Italian chub (''

Squalius cephalus ''Squalius cephalus'', the common chub, European chub or simply chub, is a species of freshwater ray-finned fish belonging to the family Leuciscidae, the daces, Eurasian minnows and related fishes. This species is found in Europe where it freq ...

''), inserting an additional "i" to prevent homonymy with the spurdog genus ''

Squalus ''Squalus'' is a genus of dogfish sharks in the family (biology), family Squalidae. Commonly known as spurdogs, these sharks are characterized by smooth dorsal fin spines, teeth in upper and lower fish jaw, jaws similar in size, caudal peduncle ...

''. In

classical Latin Classical Latin is the form of Literary Latin recognized as a Literary language, literary standard language, standard by writers of the late Roman Republic and early Roman Empire. It formed parallel to Vulgar Latin around 75 BC out of Old Latin ...

the chub and the spurdog were

homonyms In linguistics, homonyms are words which are either; ''homographs''—words that mean different things, but have the same spelling (regardless of pronunciation), or ''homophones''—words that mean different things, but have the same pronunciatio ...

as ''squalus''. An alternative explanation was that the name is a latinisation of ''squaglio'', a vernacular name for the Italian chub in Rome and its environs. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''pyrenaicus'', means "of the

Pyrenees The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...

", although this species was described from Portugal.

Distribution and habitat

''Squalius pyrenaicus'' is endemic to the catchment of the

Tagus The Tagus ( ; ; ) is the longest river in the Iberian Peninsula. The river rises in the Montes Universales between Cuenca and Teruel, in mid-eastern Spain, flows , generally westward, and empties into the Atlantic Ocean in Lisbon. Name T ...

river in Spain and Portugal. It is found in a wide variety of rivers and stream at varying altitudes and different currents, avoiding the wide, deep lowland stretches of rivers. Many of the streams it inhabits partially dry up in the summer when it uses relict pools as refuges.

Conservation

''Squalius pyrenaicus'' is classified as Vulnerable by the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...

. Threats to this species include invavsive species, pollution, droughts and habitat modification such as canalisation and damming.

References

pyrenaicus Endemic fish of the Iberian Peninsula Fish described in 1868 Taxa named by Albert Günther Taxonomy articles created by Polbot {{Leuciscinae-stub