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''Squalius'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of freshwater ray-finned fish belonging to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Leuciscidae Leuciscidae is a family of freshwater ray-finned fishes, formerly classified as a subfamily of the Cyprinidae, which contains the true minnows. Members of the Old World (OW) clade of minnows within this subfamily are known as European minnow ...
, the daces, Eurasian minnows and reklated fishes. The fishies in this genus are found in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
. Hybridization is not rare in the Leuciscidae, including this genus. '' S. alburnoides'' is known to be of ancient hybrid origin, with the paternal lineage deriving from a
prehistoric Prehistory, also called pre-literary history, is the period of human history between the first known use of stone tools by hominins  million years ago and the beginning of recorded history with the invention of writing systems. The use o ...
species related to '' Anaecypris''; the latter mated with ancestral '' S. pyrenaicus''. Present-day ''S. alburnoides'' mates with
sympatric In biology, two closely related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter each other. An initially interbreeding population that splits into two or more distinct spe ...
congeners of other
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
.Collares-Pereira, M.J. & Coelho, M.M. (2010)
Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex ''Squalius alburnoides''.
''Journal of Fish Biology, 76 (3): 707–715.''


Species

These are the currently recognized species in this genus: * '' Squalius adanaensis'' Turan, Kottelat & Doğan, 2013 (Adana chub) * '' Squalius agdamicus'' S. N. Kamensky, 1901 * '' Squalius alburnoides'' ( Steindachner, 1866) (Calandino) * '' Squalius albus'' ( Bonaparte, 1838) (Trasimeno chub) * '' Squalius anatolicus'' ( Bogutskaya, 1997) (Beyşehir chub) * '' Squalius aradensis'' ( Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) (Arade chub) * '' Squalius aristotelis'' Özuluğ & Freyhof, 2011 (Tuzla chub) * '' Squalius berak'' Heckel, 1843 (Mesopotamian chub) * '' Squalius caetobrigus'' Mendes, Perea, Sousa, Sousa-Santos & Doadrio, 2024 * '' Squalius cappadocicus'' Özuluğ & Freyhof, 2011 * '' Squalius carinus'' Özuluğ & Freyhof, 2011 (Chocolate chub) * '' Squalius carolitertii'' (Doadrio, 1988) (Northern Iberian chub) * '' Squalius castellanus'' Doadrio, Perea & Alonso, 2007 (Gallo chub) * '' Squalius cephalus'' (
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758)
(Common chub) * '' Squalius cii'' ( J. Richardson, 1857) (Marmara chub) * '' Squalius fellowesii'' ( Günther, 1868) (Aegean chub) * '' Squalius gaditanus'' Doadrio, Sousa-Santos & Perea, 2023 * '' Squalius illyricus'' Heckel & Kner, 1857 * '' Squalius irideus'' ( Ladiges, 1960) (Anatolian ghizani) * '' Squalius keadicus'' ( Stephanidis, 1971) * '' Squalius kosswigi'' ( M. S. Karaman ( sr), 1972) * '' Squalius kottelati'' Turan,
Yılmaz Yılmaz () is a Turkish language, Turkish word that translates to “unyielding”, "unbeatable", or "brave", and is a common surname and occasional male given name. Its cognates are Almas (disambiguation) and Almaz (disambiguation). Given name * ...
& Kaya, 2009
* '' Squalius laietanus'' Doadrio, Kottelat & de Sostoa, 2007 (Ebro chub) * '' Squalius latus'' Keyserling, 1861 * '' Squalius lepidus'' Heckel, 1843 (Mesopotamian pike chub) * '' Squalius lucumonis'' ( Bianco, 1983) (Etruscan chub) * '' Squalius malacitanus'' Doadrio & Carmona, 2006 (Malaga chub) * '' Squalius microlepis'' Heckel, 1843 (Imotski chub) * '' Squalius moreoticus'' (Stephanidis, 1971) (Stymphalia chub) * '' Squalius namak'' Khaefi, Esmaeili, Sayyadzadeh, Geiger & Freyhof, 2016 * '' Squalius orientalis'' ( Nordmann 1840) * '' Squalius orpheus'' Kottelat & Economidis, 2006 (Thracian chub) * '' Squalius palaciosi'' (Doadrio, 1980) (Bogardillo) * '' Squalius pamvoticus'' (Stephanidis, 1939) (Pamvotis chub) * '' Squalius peloponensis'' (
Valenciennes Valenciennes (, also , , ; ; or ; ) is a communes of France, commune in the Nord (French department), Nord Departments of France, department, Hauts-de-France, France. It lies on the Scheldt () river. Although the city and region experienced ...
, 1844)
(Peloponnese chub) * '' Squalius platyceps'' Zupančič, Marić, Naseka & Bogutskaya, 2010 * '' Squalius prespensis'' ( Fowler, 1977) (Prespa chub) * '' Squalius pursakensis'' ( Hankó ( hu), 1925) (Sakarya chub) * ''
Squalius pyrenaicus ''Squalius pyrenaicus'' , the Southern Iberian chub or Tagus chub, is a species of freshwater Actinopterygii, ray-finned fish belonging to the family (biology), family Leuciscidae, which includes the daces, Eurasian minnows and related fishes. Th ...
'' ( Günther, 1868) (Southern Iberian chub) * '' Squalius recurvirostris'' Özuluğ & Freyhof, 2011 (Akşehir chub) * '' Squalius ruffoi'' (Bianco & Recchia, 1983) * '' Squalius semae'' Turan, Kottelat & Bayçelebi, 2017 * '' Squalius seyhanensis'' Turan, Kottelat & Doğan, 2013 (Seyhan chub) * '' Squalius spurius'' Heckel, 1843 (Orontes chub) * '' Squalius squalus'' ( Bonaparte 1837) (Italian chub) * '' Squalius svallize'' Heckel & Kner, 1857 (Neretva chub) * '' Squalius tartessicus'' Doadrio, Sousa-Santos & Perea, 2023 * '' Squalius tenellus'' Heckel, 1843 (Livno chub) * '' Squalius torgalensis'' (Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) (Mira chub) * '' Squalius turcicus'' De Filippi, 1865 (Transcaucasian chub) * '' Squalius valentinus'' Doadrio & Carmona, 2006 (Eastern Iberian chub) * '' Squalius vardarensis'' S. L. Karaman, 1928 (Vardar chub) * '' Squalius verepi'' Turan, 2022 * '' Squalius zrmanjae'' S. L. Karaman, 1928 (Zrmanja chub)


References

{{Authority control Leuciscinae Taxa named by Charles Lucien Bonaparte Taxonomy articles created by Polbot