Spur-winged Lapwing
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The spur-winged lapwing or spur-winged plover (''Vanellus spinosus'') is a
lapwing Lapwings (subfamily Vanellinae) are any of various ground-nesting birds (Family (biology), family Charadriidae) akin to plovers and dotterels. They range from in length, and are noted for their slow, irregular wingbeats in flight and a shrill, ...
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
, one of a group of largish
wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
s in the family
Charadriidae The bird family Charadriidae includes the plovers, dotterels, and lapwings. The family contains 69 species that are divided into 10 genera. Taxonomy The family Charadriidae was introduced (as Charadriadæ) by the English zoologist William El ...
.


Taxonomy

The spur-winged lapwing was
formally described A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it differ ...
in 1758 by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
''. He placed it with the plovers in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Charadrius ''Charadrius'' is a genus of plovers, a group of wading birds. The genus name ''Charadrius'' is a Late Latin word for a yellowish bird mentioned in the fourth-century Vulgate. They are found throughout the world. Many ''Charadrius'' species are ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Charadrius spinosus''. He specified the type locality as Egypt. The specific epithet is from
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
meaning "thorny" (from ''spina'' meaning "thorn"). Linnaeus based his account on a description by the Swedish naturalist Fredrik Hasselquist that had been published in 1757. The spur-winged lapwing is now one of 23 species placed in the genus ''
Vanellus ''Vanellus'' is the genus of waders which provisionally contains all lapwings except red-kneed dotterel, ''Erythrogonys cinctus''. The name "''vanellus''" is Latin for "little fan", ''vanellus'' being the diminutive of ''vannus'' ("Winnowing#In ...
'' that was introduced in 1760 by the French naturalist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised.


Description

These are conspicuous and unmistakable
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s. They are medium-large waders with black crown, chest, foreneck stripe and tail. The face, the rest of the neck and belly are white and the wings and back are light brown. The bill and legs are black. Its striking appearance is supplemented by its noisy nature, with a loud ''did-he-do-it'' call. The bird's common name refers to a small
claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or Arthro ...
or spur hidden in each of its wings.


Distribution

The spur-winged lapwing breeds around the eastern
Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
, and in a wide band from sub-
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
n west
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
to
Arabia The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...
. The
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
and Turkish breeders are migratory, but other populations are resident. The species is declining in its northern range, but is abundant in much of tropical Africa, being seen at almost any wetland habitat in its range. The spur-winged lapwing is one of the species to which the ''
Agreement on the Conservation of African-Eurasian Migratory Waterbirds The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA), is an independent international treaty developed under the auspices of the United Nations Environment Programme's Convent ...
'' applies. In eastern and southern Africa the species has seen a range increase, entering
Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
for the first time in 1999 and spreading south and west.


Behaviour and ecology

This species has a preference for marshes and similar freshwater wetland habitats. The food of the spur-winged lapwing is
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s and other invertebrates, which are picked from the ground. It lays four blotchy yellowish eggs on a ground scrape. The spur-winged lapwing is known to sometimes use the wing-claws in an attack on animals and, rarely, people, who get too close to the birds' exposed offspring.


Supposed cleaning symbiosis

The "spur-winged plover" was identified by
Henry Scherren Henry James Wilson Scherren (10 February 1843 – 25 April 1911), usually known as Henry Scherren or in encyclopaedia articles as H. Sc. was the author of various books on natural history for adults and children, with notable illustrations incl ...
as the "trochilus" bird said by the Greek historian
Herodotus Herodotus (; BC) was a Greek historian and geographer from the Greek city of Halicarnassus (now Bodrum, Turkey), under Persian control in the 5th century BC, and a later citizen of Thurii in modern Calabria, Italy. He wrote the '' Histori ...
to be involved in what would now be called a
cleaning symbiosis Cleaning is the process of removing unwanted substances, such as dirt, infectious agents, and other impurities, from an object or environment. Cleaning is often performed for beauty, aesthetic, hygiene, hygienic, Function (engineering), function ...
with the
Nile crocodile The Nile crocodile (''Crocodylus niloticus'') is a large crocodilian native to freshwater habitats in Africa, where it is present in 26 countries. It is widely distributed in sub-Saharan Africa, occurring mostly in the eastern, southern, and ce ...
. However, there is no reliable evidence that this or any other species in fact has such a relationship, although Cott does record that spur-winged plovers are the birds that most often feed around basking crocodiles, and are tolerated by them.Cott, H. B. (1961). Scientific results of an inquiry into the ecology and economic status of the Nile Crocodile (''Crocodilus niloticus'') in Uganda and Northern Rhodesia. ''Transactions of the Zoological Society of London, 29'', 211-356

/ref>


Gallery

File:Vanellus spinosus egg.jpg, Egg in nest File:Vanellus spinosus MHNT.jpg, Egg - MHNT File:Spur-winged lapwing chick.jpg, Chick File:Spur-winged Lapwing (Vanellus spinosus) (21152292885).jpg, Adult File:Spur-winged lapwing (Vanellus spinosus) Gambia.jpg, A spur-winged lapwing in the
Gambia The Gambia, officially the Republic of The Gambia, is a country in West Africa. Geographically, The Gambia is the List of African countries by area, smallest country in continental Africa; it is surrounded by Senegal on all sides except for ...
File:Vanellus spinosus, Yeruham Park, israel.jpg, Underside of wings File:Spur-wingedPlover-withTongue.jpg, alt=, In Northern Israel, squawking at anyone or anything that came near its chick - showing its tongue. File:Spur-wingedPlover-withChick.jpg, alt=, In Northern Israel squawking at anyone (or anything) who came near its chick, which can be seen taking cover behind the tractor wheel.


References


External links

* * * * * * {{Taxonbar, from=Q749126 spur-winged lapwing Birds of Sub-Saharan Africa Birds of the Middle East spur-winged lapwing spur-winged lapwing