''Spinostropheus'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

of carnivorous

ceratosauria Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with '' Ceratosaurus'' than with birds. The oldest known ceratosaur, '' Saltriovenator'', dates to the earlies ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

that lived in the

Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations ...

period and has been found in the Tiouraren Formation,

Niger ) , official_languages = , languages_type = National languagestype Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * Ty ...

and only species is ''S. gautieri''.

History of discovery

In 1959,

Albert-Félix de Lapparent Albert-Félix de Lapparent (1905–1975) was a French palaeontologist. He was also a Sulpician priest. He undertook a number of fossil-hunting explorations in the Sahara desert. He contributed greatly to our knowledge of dinosaurs and other prehist ...

excavated fossils near

Oued Timmersöi Wadi ( ar, وَادِي, wādī), alternatively ''wād'' ( ar, وَاد), North African Arabic Oued, is the Arabic term traditionally referring to a valley. In some instances, it may refer to a wet ( ephemeral) riverbed that contains water o ...

, west of In Tedreft in the

Agadez Agadez ( Air Tamajeq: ⴰⴶⴰⴷⴰⵣ, ''Agadaz''), formerly spelled Agadès, is the fifth largest city in Niger, with a population of 110,497 based on the 2012 census. The capital of Agadez Region, it lies in the Sahara desert, and is a ...

desert. Among the finds were the remains of a theropod. In 1960, de Lapparent, based on these, named a second species of the genus ''

Elaphrosaurus ''Elaphrosaurus'' ( ) is a genus of ceratosaurian theropod dinosaur that lived approximately 154 to 150 million years ago during the Late Jurassic Period in what is now Tanzania in Africa. ''Elaphrosaurus'' was a medium-sized but lightly ...

'', ''E. gautieri''. The specific name honours

François Gautier François Gautier (born 1950) is a journalist based in India who served as the South Asian correspondent for multiple reputed French-language dailies. He advocates for an Indigenous Aryan narrative. He is also the founder of a private museum ...

, the discoverer of the type locality. In 2004,

Paul Sereno Paul Callistus Sereno (born October 11, 1957) is a professor of paleontology at the University of Chicago and a National Geographic "explorer-in-residence" who has discovered several new dinosaur species on several continents, including at sites ...

John Wilson John Wilson may refer to: Academics * John Wilson (mathematician) (1741–1793), English mathematician and judge * John Wilson (historian) (1799–1870), author of ''Our Israelitish Origin'' (1840), a founding text of British Israelism * John Wil ...

and

John Conrad John is a common English name and surname: * John (given name) * John (surname) John may also refer to: New Testament Works * Gospel of John, a title often shortened to John * First Epistle of John, often shortened to 1 John * Second ...

named a separate genus: ''Spinostropheus''. The generic name is derived from

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...

''spina'', "spine", and Greek στροφεύς, ''stropheus'', "vertebra", and refers to the epipophyseal processes of the cervical vertebrae, which are prominent and dorso-ventrally flattened. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

, MNHN 1961-28, was found in a layer of the Tiouraren Formation dating from the

Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. St ...

- Oxfordian. De Lapparent had presumed that the strata dated from the

Early Cretaceous The Early Cretaceous (geochronological name) or the Lower Cretaceous ( chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pr ...

. It consists of a cervical vertebra, seven pieces of the dorsals, three pieces of the sacrum, five tail vertebrae, a humerus, the lower end of a

pubic bone In vertebrates, the pubic region ( la, pubis) is the most forward-facing (ventral and anterior) of the three main regions making up the coxal bone. The left and right pubic regions are each made up of three sections, a superior ramus, inferior ra ...

, the lower end of a thighbone, a piece of a shinbone, a piece of a fibula, a metatarsal, four additional pieces of the metatarsus and a phalanx of a toe. The

paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). O ...

s were an ulna, a metatarsal and a second partial skeleton consisting of vertebrae and limb elements. In 2004, Sereno e.a. referred a third skeleton, specimen MNN TIG6 consisting of a series of cervical and dorsal vertebrae together with some ribs.

Description

''Spinostropheus'' was a relatively small theropod. In 2010,

Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...

estimated its length at 4 metres (13 feet), and its weight at 200 kg (441 lbs). In 2012 Thomas R. Holtz Jr gave a length of 6.2 meters (20.3 feet).

Phylogeny

In 2002, a

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived ch ...

analysis by Sereno ''et al'' found ''Spinostropheus'' to be the sister taxon of the Abelisauria. In this study only the data from specimen MNN TIG6 were considered. Subsequent studies have confirmed the original interpretation as a basal ceratosaur, outside of

Neoceratosauria Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with '' Ceratosaurus'' than with birds. The oldest known ceratosaur, '' Saltriovenator'', dates to the earli ...

, more closely in the evolutionary tree to ''

''. In 2018, Rafael Delcourt placed ''Spinostropheus'' in

Noasauridae Noasauridae is an extinct family of theropod dinosaurs belonging to the group Ceratosauria. They were closely related to the short-armed abelisaurids, although most noasaurids had much more traditional body types generally similar to other the ...

outside of Elaphrosaurinae and Noasaurinae.

References

{{Taxonbar, from=Q134710 Ceratosaurs Middle Jurassic dinosaurs of Africa Bathonian life Oxfordian life Mesozoic Niger Fossils of Niger Fossil taxa described in 2004 Taxa named by Paul Sereno

History of discovery

Description

Phylogeny

See also

References