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The spine-bellied sea snake (''Hydrophis hardwickii)'', also commonly known as Hardwicke's sea snake and Hardwicke's spine-bellied sea snake, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
sea snake Sea snakes, or coral reef snakes, are Elapidae, elapid snakes that inhabit Marine (ocean), marine environments for most or all of their lives. They belong to two subfamilies, Hydrophiinae and Sea krait, Laticaudinae. Hydrophiinae also includes ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Elapidae Elapidae (, commonly known as elapids , from , variant of "sea-fish") is a family of snakes characterized by their permanently erect fangs at the front of the mouth. Most elapids are venomous, with the exception of the genus '' Emydocephalus ...
. The species is native to the
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or approximately 20% of the water area of Earth#Surface, Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia (continent), ...
and the western
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...
.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''hardwickii'', is in honor of
English English usually refers to: * English language * English people English may also refer to: Culture, language and peoples * ''English'', an adjective for something of, from, or related to England * ''English'', an Amish ter ...
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
Thomas Hardwicke Major-General Thomas Hardwicke (1756 – 3 March 1835) was an English soldier and natural history, naturalist who was in India from 1777 to 1823. He collected numerous specimens of natural history and had them painted by Indian artists. From th ...
. Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Lapemis hardwickii'', p. 116).


Description

''H. hardwickii'' has the following characteristics. *Body short, stout, neck region not less than half as thick as midbody; *Head large; *
Dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
squarish or hexagonal, juxtaposed, outer 3–4 rows larger than others, scale rows: males 23–31 around neck, females 27–35, around midbody, males 25–27, females 33–41; *
Ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
small, usually distinct anteriorly, not so posteriorly; in males 114–186, in females 141–230; *Head shields entire, parietals occasionally divided; *Nostrils superior,
nasals In phonetics, a nasal, also called a nasal occlusive or nasal stop in contrast with an oral stop or nasalized consonant, is an occlusive consonant produced with a lowered velum, allowing air to escape freely through the nose. The vast majorit ...
in contact with one another; *
Prefrontal Prefrontal may refer to: *Prefrontal bone, a skull bone in some tetrapods *Prefrontal cortex, a region of the brain of a mammal *Prefrontal scales The prefrontal scales on snakes and other reptiles are the scales adjacent and anterior to the fr ...
usually in contact with second upper labial; *7–8 upper labials, 3–4 bordering eye; 1
preocular In scaled reptiles, the ocular scales are those forming the margin of the eye.Wright AH, Wright AA. 1957. ''Handbook of Snakes''. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . The name originates from the term which is Latin f ...
and 1–2 postoculars; 2, rarely 3, anterior temporals; *Greenish or yellow-olive above, whitish below; 35-50 olive to dark gray dorsal bars, tapering to a point laterally, occasionally encircling body; a narrow dark ventral stripe or broad irregular band occasionally present; *Adults often lack any pattern and are uniform olive to dark gray; *Head pale olive to black, yellow markings on snout present or not. *Total length , tail length .


Geographic range

''H. hardwickii'' is located in warm waters: *
Persian Gulf The Persian Gulf, sometimes called the Arabian Gulf, is a Mediterranean seas, mediterranean sea in West Asia. The body of water is an extension of the Arabian Sea and the larger Indian Ocean located between Iran and the Arabian Peninsula.Un ...
(
United Arab Emirates The United Arab Emirates (UAE), or simply the Emirates, is a country in West Asia, in the Middle East, at the eastern end of the Arabian Peninsula. It is a Federal monarchy, federal elective monarchy made up of Emirates of the United Arab E ...
,
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
) *Indian Ocean (
Bangladesh Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...
,
Burma Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and ha ...
,
Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
,
Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...
,
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
) *
South China Sea The South China Sea is a marginal sea of the Western Pacific Ocean. It is bounded in the north by South China, in the west by the Indochinese Peninsula, in the east by the islands of Taiwan island, Taiwan and northwestern Philippines (mainly Luz ...
north to the coasts of Fujian and Shandong * Strait of Taiwan *Indoaustralian Archipelago *North coast of
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
(
Northern Territory The Northern Territory (abbreviated as NT; known formally as the Northern Territory of Australia and informally as the Territory) is an states and territories of Australia, Australian internal territory in the central and central-northern regi ...
,
Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...
,
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
) *
Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
,
Cambodia Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
. *Pacific Ocean (
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
,
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Papua New Guinea Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
)


References


Further reading

* Boulenger GA (1896). ''Catalogue of the Snakes in the British Museum (Natural History). Volume III., Containing the Colubridæ (Opisthogylyphæ and Proteroglyphæ) ....'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiv + 727 pp. + Plates I–XXV. (''Enhydris hardwickii'', p. 301). * Gray JE (1835). ''Illustrations of Indian Zoology, chiefly selected from the collection of Major-General Hardwicke. Vol. II.'' London (1833–1834): Adolphus Richter. (Stirling, printer). 263 pp., 95 plates. (''Lapemis hardwickii'', new species, Plate 87, Figure 2). *Gray JE (1843). "Description of two new species of reptiles from the collection made during the voyages of H.M.S. Sulphur." ''Annals and Magazine of Natural History'' 'First Series'' 11: 46. * Gritis P, Voris HK (1990). "Variability and significance of parietal and ventral scales in the marine snakes of the genus ''Lapemis'' (Serpentes: Hydrophiidae), with comments on the occurrence of spiny scales in the genus." ''Fieldiana Zoology, New Series'' (56): i–iii + 1–13. * Günther ACLG (1864). ''The Reptiles of British India''. London: The Ray Society. (Taylor and Francis, printers). xxvii + 452 pp. + Plates I–XXVI. (''Hydrophis hardwickii'', p. 380 + Plate XXV, figure W). * Leviton AE, Wogan GOU, Koo MS, Zug GR, Lucas RS, Vindum JV (2003). "The Dangerously Venomous Snakes of Myanmar: Illustrated Checklist with Keys". ''Proceedings of the California Academy of Sciences'' 54 (24): 407–462. (''Lapemis hardwickii'', p. 436). * Smith MA (1943). ''The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. III.—Serpentes.'' London: Secretary of State for India. (Taylor and Francis, printers). xii + 583 pp. (''Lapemis hardwickii'', pp. 468–470, Figures 148 & 149).


External links

* SnakeDatabase (http://snakedatabase.org/species/Hydrophis/hardwickii). {{Taxonbar, from=Q2384357 Hydrophis Reptiles of Western Australia Reptiles described in 1834 Taxa named by John Edward Gray Snakes of Australia