Spicara Axillare
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''Spicara axillare'', the windtoy, is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
belonging to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Sparidae Sparidae is a family of ray-finned fishes belonging to the order Spariformes, the seabreams and porgies, although they were traditionally classified in the order Perciformes. The over 150 species are found in shallow and deep marine waters in t ...
, the seabreams and porgies. This species is found in the Western Indian Ocean and is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to South Africa.


Taxonomy

''Spicara axillare'' was first formally described as ''Caesio axillaris'' in 1900 by the Belgian-born British
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...
George Albert Boulenger George Albert Boulenger (19 October 1858 – 23 November 1937) was a Belgian-British zoologist who described and gave scientific names to over 2,000 new animal species, chiefly fish, reptiles, and amphibians. Boulenger was also an active botani ...
with its type locality given as Buffels Bay in
False Bay False Bay (Afrikaans: ''Valsbaai'') is a body of water in the Atlantic Ocean between the mountainous Cape Peninsula and the Hottentots Holland Mountains in the extreme south-west of South Africa. The mouth of the bay faces south and is demarc ...
in the
Western Cape The Western Cape ( ; , ) is a provinces of South Africa, province of South Africa, situated on the south-western coast of the country. It is the List of South African provinces by area, fourth largest of the nine provinces with an area of , an ...
of South Africa. The 5th edition of ''
Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...
'' classifies the genus ''
Spicara ''Spicara'' is a genus of ray-finned fish belonging to the family Sparidae, which includes the seabreams and porgies. These fishes are found in the Atlantic and Indian Oceans. The species in the genus are known as picarels. Taxonomy ''Spicara' ...
'' in the family Sparidae within the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Spariformes Spariformes is an order of ray-finned fishes consisting of six families within the series Percomorpha. Taxonomy Spariformes was first used as a taxonomic term in 1860 by the Dutch physician, herpetologist and ichthyologist Pieter Bleeker. Trad ...
by the 5th edition of ''
Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...
''. Some authorities classify this genus in the subfamily Boopsinae, but the 5th edition of Fishes of the World does not recognise subfamilies within the Sparidae.


Etymology

''Spicara axillare'' has the genus name ''Spicara'', which is a vernacular name for picarels, particularly ''S. flexuosa'' in Italy. This is presumed to derive from ''spica'', “a spike”, or ''spicare'', meaning “in the form of or furnished with a spike”, Rafinesque did not explain why he chose this name. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''axillare'', means
axillary Axillary means "related to the axilla (armpit)" or "related to the leaf axils". "Axillary" may refer to: Biology * Axillary artery * Axillary border * Axillary fascia * Axillary feathers * Axillary hairs * Axillary lines * Axillary lymph nodes ...
, a reference to the black spot at the base of the
pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...
.


Description

''Spicara axillare'' has its
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
supported by 11 spines and 12 soft rays while there are 3 spines and 11 soft rays supporting the
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
, there are also 17 fin rays in the pectoral fin. The pectoral and
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...
s are scythe shaped and the
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
may be crescent shaped or slightly forked. The body has a depth which fits into its
standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...
2.3 to 2.8 times. The
swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ (anatomy), organ in bony fish that functions to modulate buoyancy, and thus allowing the fish to stay at desired water depth without having to maintain lift ...
has two lobes at its rear, these lobes extend as far as the haemal spine of the 4th vertebra on the
caudal peduncle Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
. The overall colour of the body is golden, with silvery, rosy or bluish tints on the upper body. The windtoy has a maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...
of .


Distribution and habitat

''Spicara axillare'' is endemic to South Africa where it occurs from False Bay in the Western Cape to
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
. It is a
pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...
-
neritic The neritic zone (or sublittoral zone) is the relatively shallow part of the ocean above the drop-off of the continental shelf, approximately in depth. From the point of view of marine biology it forms a relatively stable and well-illuminated ...
species that is found at depths between , moving closer to shore in stormy weather.


References

{{Taxonbar, from1=Q116179777, from2=Q2182064 axillaris Taxa named by George Albert Boulenger Fish described in 1900