The speckled hind (''Epinephelus drummondhayi''), also known as the calico grouper, kitty mitchell or strawberry grouper, is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

, a

grouper Groupers are a diverse group of marine ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all members of this f ...

from the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Epinephelinae Groupers are a diverse group of marine Actinopterygii, ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all m ...

which is part of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Serranidae Serranidae is a large family (biology), family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, ...

, which also includes the anthias and sea basses. It is found in

Bermuda Bermuda is a British Overseas Territories, British Overseas Territory in the Atlantic Ocean, North Atlantic Ocean. The closest land outside the territory is in the American state of North Carolina, about to the west-northwest. Bermuda is an ...

and off the eastern coast of North America. Its natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s are open

sea A sea is a large body of salt water. There are particular seas and the sea. The sea commonly refers to the ocean, the interconnected body of seawaters that spans most of Earth. Particular seas are either marginal seas, second-order section ...

s, shallow seas, y aquatic beds, and

coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...

s. The species, like many other types of marine life in the region, is threatened by

habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...

Description

The speckled hind has a body which is robust, compressed and is deepest at the origin of the

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

, its

standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...

is 2.4 to 2.6 times its depth and is equal to the length of the head. The

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

is exposed when the mouth is closed. The margin of the gill cover bears three flat spines, while the preopercle is serrated with enlarged spines at its angle. The dorsal fin has 11 spines and 14-16 soft rays while the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

contains 3 spines and 9 soft rays. The membrane of the dorsal fin has deep indentations between the spines. The caudal fin is rounded with acutely angled corners. There are 72-76 scales in the

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

. The background colour of the head and body is dark reddish brown to grey, but it is densely speckled with tiny white spots, these sometimes merge to form a network pattern in larger individuals, the outer part of the

pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...

is yellowish. The juveniles are bright yellow and are covered in small blue-white spots. This species attains a

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

of and a maximum weight of .

Distribution

The speckled hind is found in the western Atlantic Ocean where it occurs around Bermuda as well as on the eastern coast of the United States where it is found as far north as

North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...

extending south to the

Florida Keys The Florida Keys are a coral island, coral cay archipelago off the southern coast of Florida, forming the southernmost part of the continental United States. They begin at the southeastern coast of the Florida peninsula, about south of Miami a ...

and into the

Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...

as far east as

Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...

, it also occurs off the Yucatan Peninsula. Reports of this species in the Bahamas or the

Greater Antilles The Greater Antilles is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, and Jamaica, together with Navassa Island and the Cayman Islands. Seven island states share the region of the Greater Antille ...

require confirmation.

Habitat and biology

Speckled hind adults are found on offshore areas with rocky substrates at depths of , although they are most often recorded at . The juveniles are typically found in shallower waters than the adults. Like other groupers, the speckled hind is a predatory fish and the prey taken includes other fishes,

crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...

s such as

crab Crabs are decapod crustaceans of the infraorder Brachyura (meaning "short tailed" in Greek language, Greek), which typically have a very short projecting tail-like abdomen#Arthropoda, abdomen, usually hidden entirely under the Thorax (arthropo ...

shrimp A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...

and

lobster Lobsters are Malacostraca, malacostracans Decapoda, decapod crustaceans of the family (biology), family Nephropidae or its Synonym (taxonomy), synonym Homaridae. They have long bodies with muscular tails and live in crevices or burrows on th ...

s as well as molluscs such as

squid A squid (: squid) is a mollusc with an elongated soft body, large eyes, eight cephalopod limb, arms, and two tentacles in the orders Myopsida, Oegopsida, and Bathyteuthida (though many other molluscs within the broader Neocoleoidea are also ...

. They are protogynous hermaphrodites and gather in aggregations to

spawn Spawn or spawning may refer to: * Spawning, the eggs and sperm of aquatic animals Arts, entertainment and media * Spawn (character), a fictional character in the comic series of the same name and in the associated franchise ** ''Spawn: Armageddon' ...

. The females become sexually mature at 4 or 5 years oold and at in total length and the sex change to male takes place between 7 and 14 years old. it is a long lived species which may have a life span of up to 80 years.

Taxonomy

The speckled hind was first formally described as ''Epinephelus drummondhayi'' by the American

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...

George Brown Goode George Brown Goode (February 13, 1851 – September 6, 1896), was an American ichthyologist and museum administrator. Early life and family George Brown Goode was born February 13, 1851, in New Albany, Indiana, to Francis Collier Goode and Sarah ...

(1851-1896) and

Tarleton Hoffman Bean Tarleton Hoffman Bean (October 8, 1846 – December 28, 1916) was an American ichthyologist. Biography and education Tarleton Hoffman Bean was born to George Bean and Mary Smith Bean in Bainbridge, Pennsylvania, on October 8, 1846. He attende ...

(1846-1916) in 1878 with the type locality given as

Pensacola Pensacola ( ) is a city in the Florida panhandle in the United States. It is the county seat and only city in Escambia County. The population was 54,312 at the 2020 census. It is the principal city of the Pensacola metropolitan area, which ha ...

in Florida. The specific name honours the

Scottish Scottish usually refers to something of, from, or related to Scotland, including: *Scottish Gaelic, a Celtic Goidelic language of the Indo-European language family native to Scotland *Scottish English *Scottish national identity, the Scottish ide ...

officer in the

British Army The British Army is the principal Army, land warfare force of the United Kingdom. the British Army comprises 73,847 regular full-time personnel, 4,127 Brigade of Gurkhas, Gurkhas, 25,742 Army Reserve (United Kingdom), volunteer reserve perso ...

and

ornithologist Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", is a branch of zoology dedicated to the study of birds. Several aspects of ornithology differ from related discip ...

, Lieutenant-Colonel Henry Drummond-Hay (1814-1896), originally of

Perth, Scotland Perth (; ) is a centrally located Cities of Scotland, Scottish city, on the banks of the River Tay. It is the administrative centre of Perth and Kinross council area and is the historic county town of Perthshire. It had a population of about ...

, who discovered the species on Bermuda. Since 2018 some authorities have placed this species in the genus '' Hyporthodus''

Utilisation

Speckled hind are taken as bycatch by commercial fisheries for other species of grouper and snapper off the United States coasts. It is targeted by recreational fisheries too. There is a bag limit for both commercial and recreational fisheries in the United States is one fish per vessel per trip. However, the species is still overfished and this may be because of bycatch being returned to the sea after being injured in the process of being taken.

References

{{Taxonbar, from=Q2553578 Epinephelus Fish described in 1878 Taxonomy articles created by Polbot