Southern Black Tit
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The southern black tit or simply black tit (''Melaniparus niger'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the tit family
Paridae The tits, chickadees, and titmice constitute the Paridae, a family of small passerine birds which occur mainly in the Northern Hemisphere and Africa. Many were formerly classified in the genus ''Parus''. Eurasian and African members of this f ...
, which is native to woodland habitats in southern Africa.


Subspecies

Three subspecies are recognized: * ''Melaniparus niger niger'' – southernmost
Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...
to
Eswatini Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
,
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
and
Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...
* ''Melaniparus niger xanthostomus'' – southern
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
to northern
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
, northern
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
, northwestern
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
, southern
Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
and southern
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
* ''Melaniparus niger ravidus'' – eastern Zambia,
Malawi Malawi, officially the Republic of Malawi, is a landlocked country in Southeastern Africa. It is bordered by Zambia to the west, Tanzania to the north and northeast, and Mozambique to the east, south, and southwest. Malawi spans over and ...
, Zimbabwe, Mozambique, eastern
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
and northern
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...


Range and habitat

It occurs chiefly in tropical and subtropical
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
woodland, in a semicircular arc from
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
to the
Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. It prefers woodland dominated by broad-leaved trees, most importantly
miombo Miombo woodland is a tropical and subtropical grasslands, savannas, and shrublands biome (in the World Wide Fund for Nature scheme) located in central and southern tropical Africa. It includes three woodland savanna ecoregions (listed below) chara ...
, ''
Combretum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar ...
'' species, and ''
Burkea africana ''Burkea africana'', the wild syringa (), is a deciduous, medium-sized, spreading, flat-topped tree which grows in the woodlands and savannas of sub-Saharan Africa. It is the sole species in genus ''Burkea'', which belongs to the subfamily Caesalp ...
''.''Parus niger'' (Southern Black Tit)
/ref> It is especially common in the woodlands of western
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
, where densities can reach as high as fifty birds per square kilometre.


Description

The southern black tit, unlike parids of the Northern Hemisphere or other local species, is almost entirely monotone in colour and sexually somewhat dimorphic,Newman; Kenneth;
Newman's Birds of Southern Africa
'; p. 328.
with the female being greyish and the male very distinctly blue-black in colour. It can be distinguished from the white-winged tit and Carp's tit by having no white rim in its tailHarrap, Simon; ''Tits, Nuthatches and Treecreepers''; p. 80. and is around in total length including the tail.


Food

Like all tits, it feeds chiefly on insects, and is a voracious consumer of wasps in its favoured habitat; however, it has unusually for a parid been known to take
nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...
and fruiting
fig The fig is the edible fruit of ''Ficus carica'', a species of tree or shrub in the flowering plant family Moraceae, native to the Mediterranean region, together with western and southern Asia. It has been cultivated since ancient times and i ...
s.


Breeding

Breeding occurs during the summer wet season, but its initiation is very varied and the number of broods reared depends on the consistency of rainfall. The breeding system of the southern black tit has been intensively studied and is often regarded Skutch; Alexander Frank (author) and Gardner, Dana (illustrator) ''Helpers at birds' nests: a worldwide survey of cooperative breeding and related behavior'' pp. 139-142. Published 1987 by University of Iowa Press. as a classic example of the differences in breeding strategies between warm regions with unpredictable rainfall and cold regions with severe winters. Unlike the
great tit The great tit (''Parus major'') is a small passerine bird in the tit family Paridae. It is a widespread and common species throughout Europe, the Middle East, Central Asia and east across the Palearctic to the Amur River, south to parts of No ...
or
blue tit The Eurasian blue tit (''Cyanistes caeruleus'') is a small passerine bird in the tit (bird), tit family, Paridae. It is easily recognizable by its blue and yellow plumage and small size. Eurasian blue tits, usually resident bird, resident a ...
which can provision up to twenty-five nestlings per year in unassisted pairs, the southern black tit, living on a much poorer food supply, can as an unassisted pair seldom provision even ''one'' nestling in a breeding season. As a result, most males must stay in the parental territory for several years to help rear the usually three nestlings that each breeding female produces under favourable conditions. An interesting feature of the southern black tit is that the breeding female, who incubates continuously during the fifteen days of incubation and is fed by her mate and the helpers, will
mimic In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. In the simples ...
venomous
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
s when she feels threatened to prevent depredation of the nest.


Longevity

Although the species has been recovered from
ringing Ringing may mean: Vibrations * Ringing (signal), unwanted oscillation of a signal, leading to ringing artifacts * Vibration of a harmonic oscillator ** Bell ringing * Ringing (telephony), the sound of a telephone bell * Ringing (medicine), a ri ...
on only a few occasions and data about survival and longevity are thus useless, it is practically certain that the adult mortality rate is much lower than the 70 to 75 percent mortality of closely related parids in Europe and North America.


Taxonomy

The southern black tit was formerly one of the many species in the genus ''
Parus ''Parus'' is a genus of Old World birds in the tit family Paridae. It was formerly a large genus containing most of the 50 odd species in the family Paridae. The genus was split into several resurrected genera following the publication of a deta ...
'' but was moved to ''
Melaniparus ''Melaniparus'' is a genus of birds in the tit (bird), tit family. The species were formerly placed in the speciose genus ''Parus'' but were moved to ''Melaniparus'' based on a molecular phylogenetic analysis published in 2013 that showed that the ...
'' after a
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
analysis published in 2013 showed that the members of the new genus formed a distinct
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
.


Gallery

Melaniparus niger xanthostomus, Pilanesberg NP, South Africa 3.jpg Melaniparus niger xanthostomus, Marakele NP, South Africa 4.jpg Melaniparus niger xanthostomus, Marakele NP, South Africa 1.jpg Melaniparus niger xanthostomus, Mowana Lodge, Botswana 2.jpg Southern Black Tit 2015 10 17 16 46 24 9797, crop.jpg


References


External links

* Southern Black Tit
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q27075372 southern black tit Birds of Southern Africa southern black tit Taxa named by Louis Pierre Vieillot Taxonomy articles created by Polbot