South American Land Mammal Age
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The South American land mammal ages (SALMA) establish a
geologic timescale The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochron ...
for
prehistoric Prehistory, also called pre-literary history, is the period of human history between the first known use of stone tools by hominins  million years ago and the beginning of recorded history with the invention of writing systems. The use o ...
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
n
fauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...
beginning 64.5 Ma during the
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
and continuing through to the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
(0.011 Ma). These periods are referred to as ages, stages, or intervals and were established using geographic place names where
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
materials where obtained.Flynn & Swisher, 1995 The basic unit of measurement is the first/last boundary statement. This shows that the first appearance event of one
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
is known to predate the last appearance event of another. If two taxa are found in the same fossil
quarry A quarry is a type of open-pit mining, open-pit mine in which dimension stone, rock (geology), rock, construction aggregate, riprap, sand, gravel, or slate is excavated from the ground. The operation of quarries is regulated in some juri ...
or at the same stratigraphic horizon, then their age-range zones overlap.


Background

South America was an island continent for much of the
Cenozoic The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...
, or the "Age of Mammals". As a result, its mammals evolved in their own unique directions, as Australia and Madagascar still have today.


Paleogeographic timeline

A simplified paleogeographic timeline of South America:Darin A. Croft - South American Fossil Mammals
/ref> * 66 Ma – South America was connected to both North America and Antarctica. Soon after this point, it lost its connection to North America. * 66–50 Ma – Tiupampan to Casamayoran - South America was connected to Antarctica which, in turn, was connected to Australia. The Antarctica–Australia connection was lost around the end of this interval or perhaps as much as 15 million years later. * 50–34 Ma – Casamayoran to Tinguirirican - South America was connected to Antarctica, which was not yet covered by ice. * 34 Ma –
Tinguirirican The Tinguirirican () age is a period of geologic time (36.0–29.0 Mya (unit), Ma) within the Eocene, Late Eocene and Oligocene, Early Oligocene epochs of the Paleogene, used more specifically within the South American land mammal age, SALMA classi ...
- South America and Antarctica became detached and glaciations started to form in Antarctica. * 34–9 Ma – Tinguirirican to Chasicoan - South America had no land connections to any other continent. * 9–3 Ma – Huayquerian to
Chapadmalalan The Chapadmalalan age is a period of geologic time (4.0–3.0 Ma) within the Pliocene epoch of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan and precedes the Uquian age. Fossil content ...
- islands formed between South and North America. A complete Isthmus of Panama most likely formed near the end of this interval, leading to the Great American Biotic Interchange (GABI). * 3 Ma to present –
Uquian The Uquian age is a period of geologic time (3.0–1.5 Ma) within the Pliocene and Pleistocene epochs of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan The Montehermosan age is a period of ...
to
Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
- the land connection between South and North America is established and migration between the formerly separated continents occurs. Significantly higher rates of extinction due to climatic changes occur in South America compared to North America, resulting in an asymmetrical exchange of species between the continents. North American originating taxa diversify significantly in South America during this period. It was previously believed this asymmetry was due to the main migrational route being from north to south, rather than the opposite way. * Pleistocene – the
glacial A glacier (; or ) is a persistent body of dense ice, a form of rock, that is constantly moving downhill under its own weight. A glacier forms where the accumulation of snow exceeds its ablation over many years, often centuries. It acquires ...
s and
interglacial An interglacial period (or alternatively interglacial, interglaciation) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age. The current Holocene i ...
s of the Pleistocene caused drastic eustatic sea level changes, widening and narrowing the land bridge at the 'bottleneck' of Panama. As a side-effect, the vegetation changed during this period of strong climatic changes. * Late Pleistocene – the earliest humans arrived in South America and settled in various parts of the continent. Evidence for cohabitation with the latest Pleistocene megafauna has been found at multiple locations, such as Monte Verde in coastal
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
and Tibitó on the
Altiplano Cundiboyacense The Altiplano Cundiboyacense () is a high plateau located in the Eastern Cordillera of the Colombian Andes covering parts of the departments of Cundinamarca and Boyacá. (Do not confuse with The Altiplano or the Altiplano Nariñense, both fur ...
in
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
.


Definitions


Cenozoic fossiliferous stratigraphic units in South America

The following formations have provided vertebrate, insect or plant fossils, formations with other invertebrates are excluded:


Fossil content


See also

* Biochronology ** North American land mammal age (NALMA) ** Asian land mammal age (ALMA) **
European land mammal age The European Land Mammal Mega Zones (abbreviation: ELMMZ, more commonly known as European land mammal ages or ELMA) are zones in rock layers that have a specific assemblage of fossils ( biozones) based on occurrences of fossil assemblages of Europ ...
(ELMA or ELMMZ) * List of fossiliferous stratigraphic units in Colombia * List of fossiliferous stratigraphic units in Ecuador * List of fossiliferous stratigraphic units in Paraguay * List of fossiliferous stratigraphic units in Uruguay * List of fossiliferous stratigraphic units in Venezuela


Notes and references


Notes


References


Bibliography


SALMA

* ;Paleogene * *


Pebas Wetlands

* * *


Argentina

;Paleogene * ;Pleistocene * ;Austral Basin * * ;Santa María-Hualfín Basin * * ;Santa Fe Province * ;Aconquija Formation * * * ;Agua de la Piedra Formation * * * * * ;Aisol Formation * * ;Andalhuala Formation * ;Andesitas Huancache Formation * * ;Arroyo Chasicó Formation * * * ;Belgrano Formation * ;Bororó Formation * ;Brochero Formation * * ;Casamayor Formation * * ;Cerro Azul Formation * * * ;Cerro Bandera Formation * * ;Chapadmalal Formation * * * * ;Chichinales Formation * * ;Chiquimil Formation * * * * * ;Collón Curá Formation * * * * ;Deseado Formation * * ;Divisadero Largo Formation * ;La Ensenada Formation * * * ;Las Flores Formation, Sierra del Tontal * ;Las Flores Formation, Golfo San Jorge Basin * * * ;Fortín Tres Pozos Formation * * ;Gaiman Formation * * ;Geste Formation * * * ;Gran Bajo del Gualicho Formation * ;Hernandarias Formation * ;Huayquerías Formation * * ;Huitrera Formation * ;India Muerta Formation * ;La Invernada Formation * ;Ituzaingó Formation * * * ;Koluel Kaike Formation * ;Laguna Brava Formation * ;Laguna del Hunco Formation * ;Lefipán Formation * ;Leticia Formation * ;Loma de Las Tapias Formation * * * ;Luján Formation * * ;Lumbrera Formation * * * ;Maimará Formation * * * ;Maíz Gordo Formation * ;Mariño Formation * * ;Mealla Formation * ;Miramar & San Andrés Formations * * * ;Monte Hermoso Formation * * * * * ;Monte León Formation * ;El Morterito Formation * ;Palo Pintado Formation * * * ;Paraná Formation * * * ;Peñas Coloradas Formation * * ;Pinturas Formation * * ;Piquete Formation * ;Playa de Los Lobos Allo Formation * ;Puerta del Diablo Formation * ;Puerto Madryn Formation * * ;Quebrada de Los Colorados Formation * * * * ;Río Chico Group * ;Río Foyel Formation * * ;Río Loro Formation * ;Río Mayo Formation * ;Río Negro Formation * ;Río Quinto Formation * ;Saladillo Formation * ;Salamanca Formation * * * * * ;Saldungaray Formation * ;Salicas Formation * ;Santa Cruz Formation * * * * * * * * * * * * * * * * ;Sarmiento Formation & Colhué Huapí Member * * * * * * * * * * ;Toro Negro Formation * ;Uquía Formation * * * ;Vaca Mahuida Formation * ;Ventana Formation * * ;Vorohué Formation * * * ;Yupoí Formation *


Bolivia

* * * ;Casira Formation * ;Cerdas beds * ;Honda Group * * * * ;Lacayani fauna * ;Ñuapua Formation * ;Quehua Formation * ;Salla Formation * * * ;Santa Lucía Formation * * * * * * * * * ;Tarija Formation * ;Umala Formation * ;Yecua & Petaca Formations *


Brazil

* ;Entre-Corrégos Formation * ;Fonseca Formation * ;Graxaim Formation * ;Guabirotuba Formation * ;Içá Formation * ;Itaboraí Formation * * * * * * ;Maria Farinha Formation * ;Pirabas Formation * ;Rio Madeira Formation * ;Santa Vitória Formation * ;Solimões Formation * * * * * ;Touro Passo Formation * ;Tremembé Formation * * *


Chile

;Abanico Formation * * * * * * ;Bahía Inglesa Formation * * * * * * * * * * * * * ;Caleta Herradura Formation * ;Chíu-Chíu Formation * ;Chucal Formation * ;Coquimbo Formation * ;Cura-Mallín Group * * * ;Horcón Formation * ;Huaylas Formation * * ;Loreto Formation * * * * ;Navidad Formation * ;La Portada Formation * ;Río Baguales Formation * ;Río Frías Formation * * * * *


Colombia

;Altiplano Cundiboyacense * * * * * * * * * * * * ;Cesar-Ranchería Basin * * * * * * * * * * * * * ;Cocinetas Basin * * * * * * * * * ;Barzalosa Formation * ;Gualanday Group * ;Honda Group * * * * * * * * * * * * * * * * * * * * * * * * * * * * ;Los Hoyos beds * ;Mugrosa Formation * ;Rotinet Formation * ;Sincelejo Formation *


Ecuador

;Biblián Formation * ;Dos Bocas Formation * * ;Letrero Formation * ;Onzole Formation * ;Seca Formation *


Paraguay

;Chaco Formation * *


Peru

;Peruvian Amazon * * ;Sechura Basin * ;Chambira Formation * * * ;Chilcatay Formation * * * * ;Chota Formation * ;Iñapari Formation * * ;Madre de Dios Formation * * * ;Mogollón Formation * * ;Moquegua Formation * * ;Muñani Formation * ;Paracas Formation * ;Pisco Formation * * * * * * * * * * * * * * ;Pozo Formation * * ;Soncco Formation * ;Yahuarango Formation * * * ;Yumaque Formation *


Uruguay

;Pleistocene * ;Camacho Formation * * * ;Dolores Formation * * * ;Fray Bentos Formation * * ;Raigón Formation * ;San José Formation * ;Sopas Formation *


Venezuela

;Chaguaramas Formation * ;Parángula Formation * ;Falcón Basin ;Cantaure & Paraguaná Formations * ;Capadare Formation * * ;Castillo Formation * * * * * ;Mesa Formation * ;Río Yuca Formation * ;San Gregorio Formation * * ;Santa Inés Formation * ;Urumaco, Socorro & Codore Formations * * * * *


Further reading

* * * * * * * * * * * * * {{SALMA * Regional geologic time scales