South African Cheetah
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The Southeast African cheetah (''Acinonyx jubatus jubatus'') is the nominate
cheetah The cheetah (''Acinonyx jubatus'') is a large Felidae, cat and the Fastest animals, fastest land animal. It has a tawny to creamy white or pale buff fur that is marked with evenly spaced, solid black spots. The head is small and rounded, wit ...
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
native to
East East is one of the four cardinal directions or points of the compass. It is the opposite direction from west and is the direction from which the Sun rises on the Earth. Etymology As in other languages, the word is formed from the fact that ea ...
and
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
. The Southern African cheetah lives mainly in the lowland areas and
desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
s of the
Kalahari The Kalahari Desert is a large semiarid sandy savanna in Southern Africa covering including much of Botswana as well as parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African Namib coastal d ...
, the
savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
s of Okavango Delta, and the
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s of the Transvaal region in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. In
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
, cheetahs are mostly found in farmlands. In
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
, four cheetahs of the subspecies are living in Kuno National Park in
Madhya Pradesh Madhya Pradesh (; ; ) is a state in central India. Its capital is Bhopal and the largest city is Indore, Indore. Other major cities includes Gwalior, Jabalpur, and Sagar, Madhya Pradesh, Sagar. Madhya Pradesh is the List of states and union te ...
after having been introduced there.


Taxonomy

The Southern African cheetah was first described by German naturalist
Johann Christian Daniel von Schreber Johann Christian Daniel von Schreber (17 January 1739 – 10 December 1810), often styled J.C.D. von Schreber, was a German Natural history, naturalist. Career Schreber was appointed professor of'' materia medica'' at the University of Erlangen- ...
in his book (''The Mammals illustrated as in Nature with Descriptions''), published in 1775. Schreber described the species on basis of a specimen from the
Cape of Good Hope The Cape of Good Hope ( ) is a rocky headland on the Atlantic Ocean, Atlantic coast of the Cape Peninsula in South Africa. A List of common misconceptions#Geography, common misconception is that the Cape of Good Hope is the southern tip of Afri ...
. It is therefore the
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
. Subpopulations have been called "South African cheetah" and "Namibian cheetah." Following Schreber's description, other naturalists and zoologists also described cheetah specimens from many parts of Southern and
East Africa East Africa, also known as Eastern Africa or the East of Africa, is a region at the eastern edge of the Africa, African continent, distinguished by its unique geographical, historical, and cultural landscape. Defined in varying scopes, the regi ...
that today are all considered
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
s of ''A. j. jubatus'': *''Felis guttata'' proposed in 1804 by
Johann Hermann Johann, or Jean-Frederic, Hermann, or Herrmann, (31 December 1738 in Barr, Alsace – 4 October 1800 in Strasbourg) was a French physician and naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi ...
; *''Felis fearonii'' proposed in 1834 by Andrew Smith; *''Felis lanea'' proposed in 1877 by
Philip Sclater Philip Lutley Sclater (4 November 1829 – 27 June 1913) was an England, English lawyer and zoologist. In zoology, he was an expert ornithologist, and identified the main zoogeographic regions of the world. He was Secretary of the Zoological ...
; *''Acinonyx jubatus obergi'' proposed in 1913 by Max Hilzheimer;Hilzheimer, M. (1913)
Über neue Geparden nebst Bemerkungen über die Nomenklatur dieser Tiere.
Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin: 283−292.
*''Acinonyx jubatus ngorongorensis'' proposed in 1913 by Hilzheimer on basis of a specimen from Ngorongoro,
German East Africa German East Africa (GEA; ) was a German colonial empire, German colony in the African Great Lakes region, which included present-day Burundi, Rwanda, the Tanzania mainland, and the Kionga Triangle, a small region later incorporated into Portugu ...
; *''Acinonyx jubatus velox'' proposed in 1913 by Edmund Heller on basis of a cheetah that was shot by
Kermit Roosevelt Kermit Roosevelt Sr. Military Cross, MC (October 10, 1889 – June 4, 1943) was an American businessman, soldier, explorer, and writer. A son of Theodore Roosevelt, the List of Presidents of the United States, 26th President of the United State ...
in June 1909 in the Kenyan highlands. *''Acinonyx rex'' proposed in 1927 by
Reginald Innes Pocock Reginald Innes Pocock, (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's ...
on basis of a specimen from the Umvukwe Range in
Rhodesia Rhodesia ( , ; ), officially the Republic of Rhodesia from 1970, was an unrecognised state, unrecognised state in Southern Africa that existed from 1965 to 1979. Rhodesia served as the ''de facto'' Succession of states, successor state to the ...
. In 2005, the authors of ''
Mammal Species of the World ''Mammal Species of the World: A Taxonomic and Geographic Reference'' is a standard reference work in mammalogy giving descriptions and Bibliographic database, bibliographic data for the known species of mammals. It is now in its third edition, ...
'' grouped ''A. j. guttata'', ''A. j. lanea'', ''A. j. obergi'', and ''A. j. rex'' under ''A j. jubatus'', whilst recognizing ''A. j. raineyi'' and ''A. j. velox'' as valid
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
and considering ''P. l. ngorongorensis'' as synonymous with ''raineyi''. In 2017, the Cat Classification Task Force of the Cat Specialist Group subsumed all cheetah populations from most parts of Eastern and
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
to ''A. j. jubatus'', thus making it the most widespread subspecies in the continent.


Evolutionary history

From the early
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
, the earliest African cheetah fossils have been found in the lower beds of the
Olduvai Gorge The Olduvai Gorge or Oldupai Gorge in Tanzania is one of the most important paleoanthropology, paleoanthropological localities in the world; the many sites exposed by the gorge have proven invaluable in furthering understanding of early human evo ...
site in northern Tanzania, although cheetah fossils in Southern Africa were found to be 3.5 to 3.0 million years old. The Southeast African cheetah is the second-oldest subspecies. Cheetahs from Africa and Asia were previously considered as genetically identical with each other.O’Brien S. J. 1987. East African Cheetahs: Evidence for Two Population Bottlenecks? Proceedings of the National Academy of Sciences of the United States of America. 84:508-11. DNA research and analysis started in the early 1990s and showed that the Southern and East African cheetahs are indeed separate subspecies. Until September 2009, the Asiatic cheetah was thought to be identical to African cheetahs. Stephen J. O'Brien from the Laboratory of Genomic Diversity of the
National Cancer Institute The National Cancer Institute (NCI) coordinates the United States National Cancer Program and is part of the National Institutes of Health (NIH), which is one of eleven agencies that are part of the U.S. Department of Health and Human Services. ...
was of the opinion that they have been separated from each other for only 5,000 years, which is not enough time to be classified as distinct subspecies. In early 2011, results of phylogeographic analysis of 94 cheetah samples from museum collections, wild and captive specimens, revealed that Southeast African and Asiatic cheetahs are genetically distinct, and probably diverged between 32,000 and 67,000 years ago. The
mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
data indicate that the cheetah samples from East Africa had no common
haplotype A haplotype (haploid genotype) is a group of alleles in an organism that are inherited together from a single parent. Many organisms contain genetic material (DNA) which is inherited from two parents. Normally these organisms have their DNA orga ...
with samples from Southern Africa, although one haplotype present in cheetah samples from Tanzania and Kenya clustered with samples from South Africa. The population in East Africa might have derived from a relatively recent recolonization event. The divergence between the cheetah populations in Southern, East and Northern Africa occurred between 6,700 and 32,400 years ago.


Genetics

Historically, cheetahs were thought to be genetically
homogeneous Homogeneity and heterogeneity are concepts relating to the uniformity of a substance, process or image. A homogeneous feature is uniform in composition or character (i.e., color, shape, size, weight, height, distribution, texture, language, i ...
. In 2011, results of a phylogeographic study revealed that the Asiatic cheetah and cheetah populations in Africa are genetically distinct and separated between 32,000 and 67,000 years ago. The woolly cheetah was discovered in the late 19th century by English zoologist
Philip Sclater Philip Lutley Sclater (4 November 1829 – 27 June 1913) was an England, English lawyer and zoologist. In zoology, he was an expert ornithologist, and identified the main zoogeographic regions of the world. He was Secretary of the Zoological ...
. It was considered as a separate species of cheetah that had a thicker body, and longer and denser fur. Several specimens were obtained. These creatures may be the same species as the present-day cheetah, but with a genetic disposition to long fur. In 1877, Sclater, of the
Zoological Society of London The Zoological Society of London (ZSL) is a charity and organization devoted to the worldwide animal conservation, conservation of animals and their habitat conservation, habitats. It was founded in 1826. Since 1828, it has maintained London Zo ...
, wrote of a recent acquisition by the zoo. In 1878, a second woolly cheetah was reported as a preserved specimen in the South African Museum. Both the London and South African specimens had come from Beaufort West. In 1884, a third skin was obtained from the same area, though this had more distinct spots and was a little smaller. By the late 1880s, the
trophy hunter Trophy hunting is a form of hunting for sport in which parts of the hunted wild animals are kept and displayed as trophies. The animal being targeted, known as the "game", is typically a mature male specimen from a popular species of collectable ...
s had eliminated the woolly cheetahs; from the number and locality of specimens, this variant seems to have evolved very recently (generations rather than millennia); perhaps all those animals (it seems only a handful are known at best) were the offspring of a single couple born around 1875, or maybe one more generation. The woolly cheetah has, in any case, vanished. The king cheetah was considered a different species in 1927 by naturalist
Reginald Innes Pocock Reginald Innes Pocock, (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's ...
. It was found to be a mutation caused by a recessive gene. The king cheetah is a rare variant of the Southern cheetah, first discovered in southern Rhodesia in 1925. A king cheetah was first found in South Africa in 1940 and in Botswana in 1942. However, in 1981, king cheetahs were shown to have never been a different species, as king cheetahs were born from regular parents at the De Wildt Cheetah and Wildlife Centre in South Africa, and another king cheetah were born from two female cheetahs having mated with a wild-caught male cheetah from the
Transvaal Province The Province of Transvaal (), commonly referred to as the Transvaal (; ), was a province of South Africa from 1910 until 1994, when a new constitution subdivided it following the end of apartheid. The name "Transvaal" refers to the province's ...
, and more king cheetahs were born later at the De Wildt Cheetah Centre. The king cheetahs are found in South Africa, Zimbabwe, and Botswana. In 2012, the cause of this alternative coat pattern was found to be a mutation in the gene for transmembrane aminopeptidase Q (''Taqpep''), the same gene responsible for the striped "mackerel" versus blotched "classic" patterning seen in tabby cats.. The cheetah also has
melanism Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
as one of its rare color morphs. A melanistic cheetah in Zambia was seen by Vesey Fitzgerald in the company of a spotted cheetah.


Physical characteristics

The cheetah is a medium-sized
cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
. An adult male cheetah's total size can measure from and for females. Adult cheetahs are tall at the shoulder. Males are slightly taller than females and have slightly bigger heads with wider
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s and longer
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
s. Measurements taken of wild cheetahs in Namibia indicate that the females range in head-and-body length from with long tails, and weigh between ; males range in head-and-body length from with long tails, and weigh between .. The cheetah has a bright yellow or sometimes a golden coat, and its fur is slightly thicker than that of other subspecies. The white underside is very distinct, especially on the neck and breast, and it has less spotting on its belly. The spots on the face are more pronounced, and as a whole its spots seem more dense than those of most other subspecies. The tear marks are notably thicker at the corners of the mouth, and almost all of them have distinct brown mustache markings. Like the Asiatic cheetah, it is known to have fur behind its tail and have both white and black tips at the end of its tail. However, the cheetah may also have only a black tip at the end of its tail. In desert areas, such as the
Kalahari The Kalahari Desert is a large semiarid sandy savanna in Southern Africa covering including much of Botswana as well as parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African Namib coastal d ...
, cheetahs are somewhat smaller and lighter in weight, with thinner, bright-colored fur, a trait the Northwest African cheetah also has.


Distribution and habitat

The Southeast African cheetah usually lives on
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s,
savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
s, scrub forests, and arid environments such as
desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
s and semidesert
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropica ...
s. These cheetahs can be found in open fields, where they chase and hunt herbivorous mammals such as
antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
s at a very high speed. In South Africa, the cheetah also prefers
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s (in Kruger National Park),
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
s, high mountains, mountainous grasslands, and
montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...
areas where favorable prey are mostly available. The Southeast African cheetah is currently the most common subspecies and was widespread everywhere in southern to central Africa, ranging from South Africa to the southern Democratic Republic of the Congo (
Katanga Province Katanga was one of the four large provinces created in the Belgian Congo in 1914. It was one of the eleven provinces of the Democratic Republic of the Congo between 1966 and 2015, when it was split into the Tanganyika Province, Tanganyika, Hau ...
) and southern Tanzania. Its range is now greatly reduced, where it occurs in an area of , 22% of its original range. In the past, less than 10,000 cheetahs were hunted in Namibian farmlands. Previously estimated at mere 2,000 individuals since the 1990s, as of 2015, over 3,500 cheetahs live in Namibia today. The country maintains the largest population of wild cheetahs worldwide. About 90–95% of the cheetahs live on Namibian farmlands; others live in the Kalahari Basin, the coastal deserts of
Namib The Namib ( ; ) is a coastal desert in Southern Africa. According to the broadest definition, the Namib stretches for more than along the Atlantic coasts of Angola, Namibia, and northwest South Africa, extending southward from the Carunjamba Ri ...
and Kaokoveld, and the central to northeastern region of the country. Although Namibian cheetahs are mostly found outside of
protected areas Protected areas or conservation areas are locations which receive protection because of their recognized natural or cultural values. Protected areas are those areas in which human presence or the exploitation of natural resources (e.g. firewoo ...
, they also live in Naankuse Wildlife Sanctuary,
Namib-Naukluft National Park The Namib-Naukluft National Park is a national park in western Namibia, situated between the coast of the Atlantic Ocean and the edge of the Great Escarpment. It encompasses part of the Namib Desert (considered the world's oldest desert), the ...
, and Bwabwata National Park. The cheetahs are rather uncommon in
Etosha National Park Etosha National Park is a national park in northwestern Namibia and one of the largest national parks in Africa. It was proclaimed a game reserve in March 1907 in Ordinance 88 by the Governor of German South West Africa, Friedrich von Lindequist. ...
and in Palmwag. With an approximate population of 2,000 cheetahs as of 2016, Botswana has the second-largest population of cheetahs. They are mostly found in arid habitats of the Central Kalahari, Mokolodi Nature Reserve, and
Kgalagadi Transfrontier Park Kgalagadi Transfrontier Park is a large wildlife reserve and conservation biology, conservation area in southern Africa. The park straddles the border between South Africa and Botswana and comprises two adjoining national parks: * Kalahari Gems ...
(known as Gemsbok National Park in Botswana) in the south, and in the southwest and also in the northern region of the country that holds the largest prey base, such as in Okavango Delta,
Chobe National Park Chobe National Park is Botswana's first national park, and also the most Biodiversity, biologically diverse. Located in the north of the country, it is Botswana's third largest park, after Central Kalahari Game Reserve and Gemsbok National Park. ...
, and
Moremi Game Reserve Moremi Game Reserve is a protected area in Botswana. It lies on the eastern side of the Okavango Delta and was named after Chief Moremi of the BaTawana tribe. Moremi was designated as a game reserve, rather than a national park, when it was ...
.
Khutse Game Reserve Khutse Game Reserve is a game reserve in Botswana. Overview The name Khutse means "where one kneels to drink" in Sekwena (the local dialect of Setswana). Because of its proximity, and relative accessibility, to the nation's capital, Khutse game ...
is also known to contain high abundance of suitable prey base for cheetahs, such as
springbok The springbok or springbuck (''Antidorcas marsupialis'') is an antelope found mainly in south and southwest Africa. The sole member of the genus (biology), genus ''Antidorcas'', this bovid was first Species description, described by the Germa ...
s,
gemsbok The gemsbok (''Oryx gazella''), or South African oryx, is a large antelope in the genus '' Oryx''. It is endemic to the dry and barren regions of Botswana, Namibia, South Africa and (parts of) Zimbabwe, mainly inhabiting the Kalahari and Nami ...
s, and
wildebeest Wildebeest ( , ,), also called gnu ( or ), are antelopes of the genus ''Connochaetes'' and native to Eastern and Southern Africa. They belong to the family Bovidae, which includes true antelopes, cattle, goats, sheep, and other even-toed ...
s. Cheetahs are rarely found in the eastern Botswana and at the Zimbabwean border. In South Africa, cheetahs live in the
Limpopo Limpopo () is the northernmost Provinces of South Africa, province of South Africa. It is named after the Limpopo River, which forms the province's western and northern borders. The term Limpopo is derived from Rivombo (Livombo/Lebombo), a ...
,
Mpumalanga Mpumalanga () is one of the nine provinces of South Africa. The name means "East", or literally "The Place Where the Sun Rises" in the Nguni languages. Mpumalanga lies in eastern South Africa, bordering Eswatini and Mozambique. It shares bor ...
, North West, and
Northern Cape The Northern Cape ( ; ; ) is the largest and most sparsely populated Provinces of South Africa, province of South Africa. It was created in 1994 when the Cape Province was split up. Its capital is Kimberley, South Africa, Kimberley. It includes ...
Provinces. After conservation efforts throughout the years, cheetahs have been reintroduced in the eastern, western, and southern parts, and recently in the Free State province of the country. Over 90% of the cheetah population is found outside protected areas such as
game reserve A game reserve (also known as a game park) is a large area of land where wild animals are hunted in a controlled way for sport. If hunting is prohibited, a game reserve may be considered a nature reserve; however, the focus of a game reserve ...
s and in
farmland Agricultural land is typically land ''devoted to'' agriculture, the systematic and controlled use of other forms of lifeparticularly the rearing of livestock and production of cropsto produce food for humans. It is generally synonymous with bot ...
s. More than 412 cheetahs are in
Kruger National Park Kruger National Park () is a national park in South Africa covering an area of in the provinces of Limpopo and Mpumalanga in the country's northeast. It extends from north to south and from east to west. The administrative headquarters are i ...
, subpopulations of 300 to 350 in parks and reserves, and 400 to 500 free-roaming on farmlands in the Limpopo and North West Province, although the
Kgalagadi Transfrontier Park Kgalagadi Transfrontier Park is a large wildlife reserve and conservation biology, conservation area in southern Africa. The park straddles the border between South Africa and Botswana and comprises two adjoining national parks: * Kalahari Gems ...
is the stronghold for cheetahs. Kruger and Kalahari Gemsbok National Parks hold the largest populations; they are home to roughly 42% of South Africa's cheetahs. Cheetahs had not always been common in South Africa. As of now, the country contains the third-largest population of cheetahs after years of conservation actions and reintroductions into the wild. In 2016, it is estimated about 1,500 adult cheetahs live in the wild. The population of cheetahs has been dramatically decreased in Zimbabwe, from about a thousand to 400, as of 2007. Currently, the Zimbabwean population is estimated at 165 individuals. Prior the population decline, cheetahs were more widespread in Zimbabwe, and its population had excellent growth rate, in which over 1,500 individuals thrived. Back in 1973, an estimated 400 cheetahs lived in Zimbabwe and had increased to 470 in 1987. Afterwards in 1991, a total population of 1,391 cheetahs was found by the Zimbabwe Department of Parks and Wildlife Management, whilst in 1996, a population of 728 cheetahs lived on commercial farmlands alone. In 1999, a minimum total population of 1,520 was estimated, in which over 1,200 of these cheetahs lived on commercial farmlands, while 320 were found in national parks. A year later, several reports questioned whether the Zimbabwean cheetahs were stable or decreasing, but it was increasing at the time. However, cheetahs are known to be highly threatened in farmlands in which between 1999 and 2007, 80% of the population of Zimbabwean cheetahs living in private farmlands fell into massive decline due to human-cheetah conflict, reduced from over a thousand to less than 400 as of 2007. About 100 cheetahs were killed by livestock farmers in Zimbabwe's lowveld per year. Following years later, about 150 to 170 adult cheetahs are found there, and the human-cheetah conflict is no longer a major threat to the species. Most of the Zimbabwean cheetahs live in protected areas today. Cheetahs are mainly found in the southern to central regions. Isolated populations are found in northwestern Zimbabwe, such as
Victoria Falls Victoria Falls (Lozi language, Lozi: ''Mosi-oa-Tunya'', "Thundering Smoke/Smoke that Rises"; Tonga language (Zambia and Zimbabwe), Tonga: ''Shungu Namutitima'', "Boiling Water") is a waterfall on the Zambezi River, located on the border betwe ...
, Matetsi, and Kazuma Pan, also near the Mozambican border.
Hwange National Park Hwange National Park (formerly Wankie Game Reserve) is the largest natural reserve in Zimbabwe. It is around 14,600 sq km in area. It lies in the northwest of the country, just off the main road between Bulawayo and Victoria Falls. The nearest ...
, the largest reserve with an area of , is the main stronghold for the Zimbabwean cheetahs. Cheetahs also live in Matobo National Park. The cheetahs of the Zambezi Valley are nearly extinct, as only three individuals remain in
Matusadona National Park Matusadona National Park is a national park in northern Zimbabwe situated on the southern shore of Lake Kariba. The park takes its name from the local Matuzviadonha Hills. History Matusadona was proclaimed a non-hunting area on 7 November 1958 ...
and 9 in the
Mana Pools National Park Mana Pools National Park is a wildlife conservation area and national park in northern Zimbabwe.lowveld Veld ( or , Afrikaans and Dutch: ''veld'', field), also spelled veldt, is a type of wide-open, rural landscape in Southern Africa. Particularly, it is a flat area covered in grass or low scrub, especially in the countries of South Africa, ...
, most of which live in Gonarezhou National Park, private reserves (Bubye, Save, Malilangwe, Nuanetsi), and at the Chilojo Cliffs. In Zambia, cheetahs are mostly spotted at Matamene Camp of
Liuwa Plain National Park Liuwa Plain National Park is a national park in Zambia's Western Province, Zambia, Western Province. "Liuwa" means "plain" in the local Liuwa language, a dialect of Lozi language, and the plains originally served as a hunting ground for Lewa ...
from the Western Province. The national park is part of the Kavango–Zambezi Transfrontier Conservation Area. They are also present at the Kafue National Park, near the
Kafue River The long Kafue River is the longest river lying wholly within Zambia. Its water is used for irrigation and for generating Hydroelectricity, hydroelectric power. It is the largest tributary of the Zambezi, and of Zambia's principal rivers, it ...
and at the Sioma Ngwezi National Park (the second-largest park in Africa) in the southwest corner of Zambia. About 100 cheetahs live in the country. In 2007, between 50 and 90 cheetahs were estimated to survive in Mozambique, where the species inhabits grasslands, savannahs, and mixed ''
Acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
'' and
mopane ''Colophospermum mopane'', commonly called mopane, mopani, butterfly tree, turpentine tree, or balsam tree, is a tree in the legume family (Fabaceae), that grows in hot, dry, low-lying areas, in elevation, in parts of Southern Africa. The tree ...
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s. Most habitats consist of
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
s and rivers. Historically, it was widespread in the country, but by 1975, the population had declined to about 200 individuals due to intense poaching during the
Mozambican Civil War The Mozambican Civil War () was a civil war fought in Mozambique from 1977 to 1992 due to a combination of local strife and the polarizing effects of Cold War politics. The fighting was between Mozambique's ruling Marxist Front for the Liberat ...
. Camera traps set up in 2004 and 2011 revealed constant presence of cheetahs, other predators and herbivores in Mozambique's conservation areas in
Great Limpopo Transfrontier Park Great Limpopo Transfrontier Park is a 35,000 km2 peace park that is in the process of being formed. It will link the Limpopo National Park (formerly known as Coutada 16) in Mozambique, Kruger National Park in South Africa, Gonarezhou Natio ...
. Around 35 cheetahs live in
Limpopo National Park The Limpopo National Park () was born when the status of Coutada 16 Wildlife Utilisation Area in Gaza Province, Mozambique, was changed from a hunting concession to a protected area. It forms part of the Great Limpopo Transfrontier Park with the ...
. Cheetahs are also present in Zinave National Park and Banhine National Park, which are part of the Great Limpopo Transfrontier Park. Apart from the central and northern regions, the now-rare cheetah lived in southern Tanzania. It ranged close to the Zambia/Malawi borders in the southwest to the southeasternmost part of the country. These cheetahs are found at Mpanga-Kipengere Game Reserve and the Uwanda Game Reserve. Whether or not they are extinct in
Selous Game Reserve The Selous Game Reserve, now renamed as Nyerere National Park (in-part), is a protected nature reserve and wilderness area in southern Tanzania, East Africa. It covers a total area of , with additional buffer zones, as well. It was designated a ...
is unknown. The indigenous population of cheetahs was extinct in
Eswatini Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
. In 1997, three cheetahs have been reintroduced into the
Hlane Royal National Park Hlane Royal National Park is a national park in Eswatini, roughly 67 km northeast of Manzini along the MR3 road.South Africa, page 815 Prior to being designated a public park, it was a private royal hunting ground. Hlane, meaning 'wilderne ...
, the largest () protected area of Eswatini. The cheetah was once thought to be extinct in Angola, but in 2010 two adult male cheetahs were spotted in the
Iona National Park Iona National Park ( Portuguese: ) is the largest and oldest national park in Angola. It is situated in the Southwestern corner of the country, in Namibe Province. It is roughly bound by the Atlantic Ocean to the West, an escarpment to the East ...
. It was the first time cheetahs have been sighted in the wild of Angola in 30 years. This protected area provides suitable habitat for the cheetah, as it has a large, open savannah where springbok and
oryx ''Oryx'' ( ) is a genus consisting of four large antelope species called oryxes. Their pelage is pale with contrasting dark markings in the face and on the legs, and their long horns are almost straight and annulated. The exception is the sci ...
occur. In the 1980s, cheetahs occurred in three protected areas, namely
Kasungu National Park Kasungu National Park is a national park in Malawi. It is located west of Kasungu, about 175 km north of Lilongwe, extending along the Zambian border. Kasungu National Park, established in 1970, is the second-largest in Malawi at , with an ...
,
Nyika National Park Nyika National Park is Malawi’s largest national park, with an area of 3200 km2 (1250 mile2). Location and road access The park covers practically the whole of the Nyika Plateau in northern Malawi, about 480 km north of Lilongwe and 60 km no ...
and Vwaza Marsh Game Reserve, also at the Zambian border in the west. In 2007, the small cheetah population was considered locally extinct due to loss of suitable habitat and lack of prey.


Former range

In early 20th century, African cheetahs were widespread everywhere in the continent, until they lost most of their ranges and disappeared from 23 countries. Cheetahs are locally extinct from
Lesotho Lesotho, formally the Kingdom of Lesotho and formerly known as Basutoland, is a landlocked country in Southern Africa. Entirely surrounded by South Africa, it is the largest of only three sovereign enclave and exclave, enclaves in the world, t ...
and the southern part of the
Democratic Republic of the Congo The Democratic Republic of the Congo (DRC), also known as the DR Congo, Congo-Kinshasa, or simply the Congo (the last ambiguously also referring to the neighbouring Republic of the Congo), is a country in Central Africa. By land area, it is t ...
. They used to live in the Katanga, southern Bandundu, southern Kasai-Occidental, and
Bas-Congo Kongo Central (), formerly Bas-Congo, is one of the 26 provinces of the Democratic Republic of the Congo. Its capital is Matadi. History At the time of independence, the area now encompassing Kongo Central was part of the greater province of ...
. They probably extended to
Kinshasa Kinshasa (; ; ), formerly named Léopoldville from 1881–1966 (), is the Capital city, capital and Cities of the Democratic Republic of the Congo, largest city of the Democratic Republic of the Congo. Kinshasa is one of the world's fastest-grow ...
. It was previously estimated that about 100 to 500 cheetahs lived in the Congo between 1950-70s. Kundelungu National Park was one of the remaining stronghold for the cheetahs after recent extinction of cheetahs from
Virunga National Park Virunga National Park is a national park in the Albertine Rift Valley in the eastern part of the Democratic Republic of the Congo. It was created in 1925. In elevation, it ranges from in the Semliki River valley to in the Rwenzori Mountains. ...
and Sudan cheetahs from
Garamba National Park Garamba National Park () is a national park in the north-eastern Democratic Republic of the Congo covering nearly . It is among Africa's oldest parks and was designated a World Heritage Site by UNESCO in 1980 for its protection of critical habit ...
during the 1960s. Rumored sightings have been reported occasionally in the southern regions of the Congo, such as near the Angolan border, around the Sandoa Territory and on the Kibara Plateau of
Upemba National Park Upemba National Park ( French: ''Parc national d'Upemba'') is a large national park in Haut-Lomami, Lualaba Province & Haut-Katanga Province (formerly in Katanga Province) of the southeastern Democratic Republic of Congo, formerly Zaire. Geogr ...
. To this day, Upemba is still considered as the only national park of the country to contain cheetah populations.


Ecology and behavior


Reproduction and lifecycle

Male cheetahs are sociable and may live in a group with other male siblings. Males establish their territories by marking their territories by urinating on trees or termite mounds. The females, though, are not sociable and do not establish a territory. They are solitary and avoid each other. However, they may live with their mothers, daughters, or sisters on their home ranges. The female's home range's size can depend on the prey base. Cheetahs in southern African
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s have ranges as small as , while in some parts of Namibia, they can reach . Female cheetahs can reproduce at 13 to 16 months of age and with a typical age of sexual maturity between 20 and 23 months. The gestation can last for 90 to 95 days. Cub births mostly occur at November to January in Namibia and November to March in Zambia. Females hunt solo, except the cheetah cubs accompany their mothers to learn how to hunt on their own after the age of 5–6 weeks. After the cubs reach 18 months of age, the mother leaves her cubs, and the siblings remain as a group for a few months until the sisters leave the group and the brothers stay together. The male cubs may form alliances with other males after separating from their mother.


Hunting and diet

The cheetah is a carnivorous mammal. It preys on medium-sized and large
antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
s, and fast, small animals including
Cape hare The Cape hare (''Lepus capensis''), also called the brown hare and the desert hare, is a hare native to Africa and Arabia extending into India. Taxonomy The Cape hare was one of the many Mammalia in the 10th edition of Systema Naturae, mammal ...
s and rodents. It prefers
Thomson's gazelle Thomson's gazelle (''Eudorcas thomsonii'') is one of the best known species of gazelles. It is named after explorer Joseph Thomson (explorer), Joseph Thomson and is sometimes referred to as a "tommie". It is considered by some to be a subspecies o ...
,
impala The impala or rooibok (''Aepyceros melampus'', lit. 'black-footed high-horn' in Ancient Greek) is a medium-sized antelope found in eastern and southern Africa. The only extant member of the genus '' Aepyceros'', and tribe Aepycerotini, it ...
,
kudu The kudus are two species of antelope of the genus '' Tragelaphus'': * Lesser kudu, ''Tragelaphus imberbis'', of eastern Africa * Greater kudu, ''Tragelaphus strepsiceros'', of eastern and southern Africa The two species look similar, th ...
,
puku The puku (''Kobus vardonii'') is a medium-sized antelope found in wet grasslands in southern Democratic Republic of Congo, Namibia, Tanzania, Zambia and more concentrated in the Okavango Delta in Botswana. Nearly one-third of all puku are found ...
, oribi,
springbok The springbok or springbuck (''Antidorcas marsupialis'') is an antelope found mainly in south and southwest Africa. The sole member of the genus (biology), genus ''Antidorcas'', this bovid was first Species description, described by the Germa ...
,
gemsbok The gemsbok (''Oryx gazella''), or South African oryx, is a large antelope in the genus '' Oryx''. It is endemic to the dry and barren regions of Botswana, Namibia, South Africa and (parts of) Zimbabwe, mainly inhabiting the Kalahari and Nami ...
,
steenbok The steenbok (''Raphicerus campestris'') is a common small antelope of southern and eastern Africa. It is sometimes known as the steinbuck or steinbok. Description Steenbok resemble small oribi, standing 45–60 cm (16"–24") at the ...
,
wildebeest Wildebeest ( , ,), also called gnu ( or ), are antelopes of the genus ''Connochaetes'' and native to Eastern and Southern Africa. They belong to the family Bovidae, which includes true antelopes, cattle, goats, sheep, and other even-toed ...
,
warthog ''Phacochoerus'' is a genus in the family Suidae, commonly known as warthogs (pronounced ''wart-hog''). They are pigs who live in open and semi-open habitats, even in quite arid regions, in sub-Saharan Africa. The two species were formerly cons ...
,
red hartebeest The red hartebeest (''Alcelaphus buselaphus caama''), also called the Cape hartebeest or Caama, is a subspecies of the hartebeest found in Southern Africa. More than 130,000 individuals live in the wild. The red hartebeest is closely related to ...
,
nyala The lowland nyala or simply nyala (''Tragelaphus angasii'') is a spiral-horned artiodactyl antelope native to Southern Africa. The species is part of the family Bovidae and the genus '' Tragelaphus'' (formerly placed in the genus ''Nyala''). It ...
and other
ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
s.


Enemies and competitors

Like other cheetahs, they are threatened and outranked by larger predators in their area. They are threatened by
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...
s,
leopard The leopard (''Panthera pardus'') is one of the five extant cat species in the genus ''Panthera''. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of with a ...
s, painted dogs,and
hyena Hyenas or hyaenas ( ; from Ancient Greek , ) are feliform carnivoran mammals belonging to the family Hyaenidae (). With just four extant species (each in its own genus), it is the fifth-smallest family in the order Carnivora and one of the sma ...
s as they can steal their carcasses and kill them if given the chance. The cheetahs surrender their meals to
spotted hyena The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...
s. However, coalitions of male adult cheetahs can chase predators away, and a single cheetah can chase
jackal Jackals are Canidae, canids native to Africa and Eurasia. While the word has historically been used for many canines of the subtribe Canina (subtribe), canina, in modern use it most commonly refers to three species: the closely related black-b ...
s and lone
wild dog A free-ranging dog is a dog that is not confined to a yard or house. Free-ranging dogs include street dogs, village dogs, stray dogs, feral dogs, etc., and may be owned or unowned. The global dog population is estimated to be 900 million, of whi ...
s away.


Threats

The Southern cheetah is a vulnerable subspecies, due to
poaching Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...
,
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
, and lack of prey. Indiscriminate capture and removal of wild cheetahs in southern Africa continue to threaten the survival of this species, as it may reduce the
genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species. It ranges widely, from the number of species to differences within species, and can be correlated to the span of survival for a species. It is d ...
in the wild and they breed poorly in captivity. Its survival is also threatened by
inbreeding Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely genetic distance, related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genet ...
. In
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
, the cheetahs are mostly threatened by habitat changes. The cheetah was also highly threatened by
hunting Hunting is the Human activity, human practice of seeking, pursuing, capturing, and killing wildlife or feral animals. The most common reasons for humans to hunt are to obtain the animal's body for meat and useful animal products (fur/hide (sk ...
and range loss. In early 1930s, the cheetahs were hunted down and almost went extinct in South Africa. Therefore, it has lost most of its range, mostly in South Africa and Mozambique. Only a few dozen of them live in the southern part of Mozambique. It also disappeared from many regions of South Africa, only living in the northern and northeastern parts of the country. During the 1970s, 9,500 cheetahs were killed in Namibian farmlands. As a protected species in Namibia, people are allowed to remove Namibian cheetahs only if they pose a threat to livestock or human life. Unfortunately, farmers might capture Namibian cheetahs, often removing or killing those that have not taken any livestock. About 90% of the Namibian cheetah population live on farmlands. In Botswana, the cheetah is protected under the Conserved Animal legislation since 1968, which strictly limits hunting and capture. Before then, the decline of suitable prey caused the cheetahs to feed on livestock. About 50 cheetah were previously hunted down by tribesmen each year to protect livestock. Limited international trade in live animals and skins is permitted from Namibia, Zimbabwe, and Botswana.


Conservation status

Previously estimated at a population of 4,190 individuals in Southern Africa since 2007, the total population of the Southern cheetah has likely reached over 6,000 individuals,IUCN/SSC. (2007). Regional conservation strategy for the cheetah and African wild dog in Southern Africa. IUCN Gland, Switzerland. with Namibia having the largest cheetah population worldwide. Since 1990, the population was estimated to be about 2,500 individuals in Namibia; until 2015, the cheetah population has increased to more than 3,500 in the country. Botswana contains the second-largest population of cheetahs – in 2007, an estimated population of 1,800. However, in 2016, about 2,000 cheetahs were in Botswana, which is about 20% of the world's cheetahs. Around 550 to 850 cheetahs were left in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
in 2007. After many conservation efforts, the cheetah population has boosted to more than 1,000. In 2013, the estimated population was between 1,200 and 1,300 cheetahs in South Africa. Whilst an estimated that 1,500 adult cheetahs live in South Africa since 2016, the Endangered Wildlife Trust stated that the total population ranges between 1,166 and 1,742 cheetahs in South Africa alone in 2017. In
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
, on the contrary, the cheetahs' population has severely declined, from more than 1,500 cheetahs since 1999 to 400 cheetahs in 2007, to between 150 and 170 cheetahs as of 2015. In 2007, around 100 individuals remained in
Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
and between 50 and 90 were left in
Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...
. Several conservation projects for the cheetah species exist in African countries and
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
. Like the Asiatic cheetah, the Southern cheetah got more attention from people than other subspecies. Three cheetah subspecies are included on the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
list of
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
(three African subspecies threatened, Northwest African and Asiatic subspecies in critical condition), as well as on the US
Endangered Species Act The Endangered Species Act of 1973 (ESA; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting and conserving imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of e ...
:
threatened species A threatened species is any species (including animals, plants and fungi) which is vulnerable to extinction in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensatio ...
- Appendix I of
CITES CITES (shorter acronym for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of inte ...
(Convention on International Trade in Endangered Species). Founded in Namibia in 1990, the Cheetah Conservation Fund's mission is to be the world's resource charged with protecting the cheetah and to ensure its future. The organization works with all stakeholders within the cheetah's ecosystem to develop best practices in research, education, and ecology, and create a sustainable model from which all other species, including people, will benefit. Around 12,400 cheetahs were estimated to remain in the wild in 25 African countries. Recently, 6,674 mature individuals were found by the IUCN; Namibia has the most, with more than 3,500, of which 90% of them are living outside of protected areas. Breeding programs have been successful, including the use of ''in vitro'' fertilization, in zoos around the world. The cheetahs are known to be poor breeders in captivity, though several organizations, such as the De Wildt Cheetah and Wildlife Centre, have succeeded in breeding high numbers of cheetah cubs. In 2009, the centre has bred more than 800 cubs.


Reintroduction projects


In Africa


South Africa

The cheetah once occurred in several areas of
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
, but not commonly in South Africa. The species lives mostly on the eastern and northern locations of South Africa. Since the 1960s, the cheetah had been imported from Namibia, which used to contain healthy populations of cheetahs at the time, and has been reintroduced to their former ranges and in small reserves. About 29% of the cheetah population was indigenously from South Africa whilst 71% was those imported from Namibia. The first known reintroductions were in
KwaZulu Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
,
Gauteng Gauteng ( , ; Sotho-Tswana languages, Sotho-Tswana for 'place of gold'; or ) is one of the nine provinces of South Africa. Situated on the Highveld, Gauteng is the smallest province by land area in South Africa. Although Gauteng accounts f ...
,
Lowveld Veld ( or , Afrikaans and Dutch: ''veld'', field), also spelled veldt, is a type of wide-open, rural landscape in Southern Africa. Particularly, it is a flat area covered in grass or low scrub, especially in the countries of South Africa, ...
,
Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...
,
Western Cape The Western Cape ( ; , ) is a provinces of South Africa, province of South Africa, situated on the south-western coast of the country. It is the List of South African provinces by area, fourth largest of the nine provinces with an area of , an ...
, and Southern Kalahari. There are currently 1,500 mature cheetahs within the country. In December 2003, after the cheetahs were heavily hunted in the
Great Karoo The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe Khoemana (also known as !Orakobab or Korana) word is a semidesert natural region of South Africa. No exact definition of what constitutes the Karoo is available, so its extent i ...
and Eastern Cape areas to extinction 125 years ago, cheetahs have returned to the
Karoo The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe Khoemana (also known as !Orakobab or Korana) word is a semidesert natural region of South Africa. No exact definition of what constitutes the Karoo is available, so its extent is ...
, starting with a severely injured female wild-born cheetah named Sibella (''circa'' 2001 – 11 September 2015) that went through surgery at the De Wildt Cheetah and Wildlife Trust and was later reintroduced to Samara Private Game Reserve. The reintroduction process was a success. Sibella was a capable hunter and successfully raised 18 cubs. About 2% of the wild population of cheetahs in South Africa occurs in that region. The cheetahs living in Samara are also in better condition, threats from
apex predator An apex predator, also known as a top predator or superpredator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the hig ...
s such as lions and hyenas no longer exist. Sibella's youngest daughter Chilli has given birth to the first third-generation cheetah cubs of the Samara Private Game Reserve in January 2017. A National Cheetah Metapopulation Project was launched in 2011 by the Endangered Wildlife Trust. Its purpose is to develop and co-ordinate a national
metapopulation A metapopulation consists of a group of spatially separated populations of the same species which interact at some level. The term metapopulation was coined by Richard Levins in 1969 to describe a model of population dynamics of insect pests in a ...
management plan for cheetahs in smaller fenced reserves in South Africa. For instance, the cheetahs have been reintroduced in around 50 of these South African reserves. Fragmented subpopulations of cheetahs are currently increasing in a few hundreds. As of July 2014, further plans exist to reintroduce the cheetahs in six more small fenced reserves over the next few years. For the first time after 100 years of extinction since the colonial period, the cheetah has recently been reintroduced into the Free State in 2013, with two male wild cheetahs that have been relocated from the
Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...
's Amakhala Game Reserve to the Free State's Laohu Valley Reserve, where the
critically endangered An IUCN Red List critically endangered (CR or sometimes CE) species is one that has been categorized by the International Union for Conservation of Nature as facing an extremely high risk of extinction in the wild. As of December 2023, of t ...
South China tiger from
Save China's Tigers Save China's Tigers (SCT) is an international charitable foundation based in Hong Kong, the United States, and the United Kingdom (headquartered in London) that aims to save the big cats of China from extinction. It focuses on the Chinese tigers ...
(SCT) are part of a
rewilding Rewilding is a form of ecological restoration aimed at increasing biodiversity and restoring natural processes. It differs from other forms of ecological restoration in that rewilding aspires to reduce human influence on ecosystems. It is also d ...
project in South Africa. A female cheetah has yet to be reintroduced to Laohu Valley. In early 2016, an adult female has been reintroduced to the reserve. Three wild cheetah cubs have been born for the first time in Laohu Valley Reserve in February 2017, making them the first cheetahs born in the wild since their disappearance from the Free State province in over a century. With three mature individuals and three cubs, six cheetahs are in Laohu Valley Reserve. In 2016, a reintroduction and rewilding project known as Rewilding iSimangaliso for cheetahs is going on in iSimangaliso situated in
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
province, including
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...
s and Cape wild dogs, which were first introduced in late 2013. The 15 collared resident cheetahs in uMkhuze Game Reserve are seen by visitors during game drives.


Malawi

In May 2017, two male and two female cheetahs were imported from South Africa and reintroduced to
Liwonde National Park Liwonde National Park, also known as Liwonde Wildlife Reserve, is a national park in southern Malawi, near the Mozambique border. The park was established in 1973, and has been managed by the nonprofit conservation organization African Parks since ...
.


Zambia

Since 1989, only a few cheetahs had been recorded at the Lower Zambezi National Park, despite the area being apparently suitable habitat for cheetahs. Chiawa Camp, in association with National Parks and Wildlife and Japan Aid, approached the Cheetah Conservation Fund for a study group to assess the suitability of Lower Zambezi. In October 1994, reintroduction attempts were made with three cheetahs to the Lower Zambezi. However, the reintroduction project had been unsuccessful, as two of them were killed by traps; one survivor remained for three years alone. Further plans exist to reintroduce the cheetah to the Lower Zambezi.


Outside Africa

Asiatic cheetahs had existed in
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
for thousands of years, but as a result of hunting and other disastrous causes, they have disappeared there, with the last known Indian cheetah having been spotted in 1951. The critically endangered species currently lives in Iran, as the country itself is unwilling to give their cheetahs to India. A captive propagation project has been proposed. Minister of Environment and Forests
Jairam Ramesh Jairam Ramesh (born 9 April 1954) is an Indian politician belonging to the Indian National Congress. He is a Member of Parliament representing Karnataka state in the Rajya Sabha. In July 2011, Jairam was elevated to the Union Council of Minis ...
told the
Rajya Sabha Rajya Sabha (Council of States) is the upper house of the Parliament of India and functions as the institutional representation of India’s federal units — the states and union territories.https://rajyasabha.nic.in/ It is a key component o ...
on 7 July 2009, "The cheetah is the only animal that has been described extinct in India in the last 100 years. We have to get them from abroad to repopulate the species." He was responding to a call for attention from Rajiv Pratap Rudy of the
Bharatiya Janata Party The Bharatiya Janata Party (BJP; , ) is a political party in India and one of the two major List of political parties in India, Indian political parties alongside the Indian National Congress. BJP emerged out from Syama Prasad Mukherjee's ...
(BJP). "The plan to bring back the cheetah, which fell to indiscriminate
hunting Hunting is the Human activity, human practice of seeking, pursuing, capturing, and killing wildlife or feral animals. The most common reasons for humans to hunt are to obtain the animal's body for meat and useful animal products (fur/hide (sk ...
and complex factors like a fragile
breeding Breeding is sexual reproduction that produces offspring, usually animals or plants. It can only occur between a male and a female animal or plant. Breeding may refer to: * Animal husbandry, through selected specimens such as dogs, horses, and rab ...
pattern is audacious given the problems besetting tiger conservation." Two naturalists, Divya Bhanusinh and MK Ranjit Singh, suggested importing cheetahs from Namibia, after which they will be bred in captivity and, in time, released in the wild. Multiple suitable potential sites from the Indian states of
Gujarat Gujarat () is a States of India, state along the Western India, western coast of India. Its coastline of about is the longest in the country, most of which lies on the Kathiawar peninsula. Gujarat is the List of states and union territories ...
,
Madhya Pradesh Madhya Pradesh (; ; ) is a state in central India. Its capital is Bhopal and the largest city is Indore, Indore. Other major cities includes Gwalior, Jabalpur, and Sagar, Madhya Pradesh, Sagar. Madhya Pradesh is the List of states and union te ...
, and
Rajasthan Rajasthan (; Literal translation, lit. 'Land of Kings') is a States and union territories of India, state in northwestern India. It covers or 10.4 per cent of India's total geographical area. It is the List of states and union territories of ...
which consisted of forests, grasslands, savannahs, and deserts were chosen for the cheetah reintroduction project in India, such as
Banni Grasslands Reserve Banni Grasslands Reserve or Banni grasslands form a belt of arid grassland ecosystem on the outer southern edge of the desert of the marshy salt flats of Rann of Kutch in Kutch District, Gujarat State, India. They are known for rich wildlife a ...
, Desert National Park, Kuno National Park and Gajner Wildlife Sanctuary. They are also known to be where the Asiatic cheetahs and other native animals coexisted for several years until they had recently gone extinct from the region. Nauradehi Wildlife Sanctuary was chosen as the most suitable site for reintroduced Namibian cheetahs. However, the plan to introduce this subspecies to India has been suspended in 2012, after discovering the distinctness between the cheetahs from Asia and Africa, having been separated between 32,000 and 67,000 years ago. In 2020, the
Supreme Court of India The Supreme Court of India is the supreme judiciary of India, judicial authority and the supreme court, highest court of the Republic of India. It is the final Appellate court, court of appeal for all civil and criminal cases in India. It also ...
ruled that African cheetahs, a different subspecies, could be brought into the country at a "carefully chosen location" on an experimental basis. In 2022, India received eight cheetahs from Namibia last year and they were released in Kuno National Park in
Madhya Pradesh Madhya Pradesh (; ; ) is a state in central India. Its capital is Bhopal and the largest city is Indore, Indore. Other major cities includes Gwalior, Jabalpur, and Sagar, Madhya Pradesh, Sagar. Madhya Pradesh is the List of states and union te ...
state. In 2023, South Africa signed an agreement with India to introduce 12 African cheetahs to India each year over the next decade. As of 22 March 2023, four African cheetahs were living in the wild of Kuno National Park in Madhya Pradesh.


In captivity

Cheetahs are known to be difficult to breed in captivity because of their social behaviors and breeding problems. The cub mortality in captivity and in the wild is high at about 50%. On average, 30% of all captive-bred cubs born in captivity may die within a month. The Southern cheetah is the most widespread subspecies breeding in captivity around the world, while Sudanese cheetahs are found only in a few European and
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
ern zoos and wildlife centers. The subspecies is found in various zoos worldwide in America, Africa, Eurasia, and Australia. Several zoos, facilities, breeding centers, and wildlife parks part of the American (
Species Survival Plan The American Species Survival Plan or SSP program was developed in 1981 by the (American) Association of Zoos and Aquariums to help ensure the survival of selected species in zoos and aquariums, most of which are threatened or endangered in the w ...
,
Association of Zoos and Aquariums The Association of Zoos and Aquariums (AZA), originally the American Association of Zoological Parks and Aquariums (AAZPA), is an American 501(c)(3) nonprofit organization founded in 1924 and dedicated to the advancement of zoos and public aqu ...
) and Eurasian (
European Endangered Species Programme The EAZA Ex-situ Programme (EEP) is a population management and Ex situ conservation, conservation programme by European Association of Zoos and Aquaria, European Association of Zoos and Aquaria (EAZA) for wild animals living in European zoos. The p ...
,
European Association of Zoos and Aquaria The European Association of Zoos and Aquaria (EAZA) is an organisation for the European zoo and aquarium community that links over 340 member organisations in 41 countries. EAZA membership is open to all zoos and aquaria across Europe that compl ...
) captive-breeding programs have been successfully increasing populations of cheetahs, such as
White Oak Conservation White Oak Conservation, which is part of Walter Conservation, is a conservation center in northeastern Florida. It is dedicated to the conservation of Endangered species, endangered and threatened species, including Indian rhinoceros, southern ...
from
Yulee, Florida Yulee is a census-designated place (CDP) located within Nassau County, Florida, in the United States. Its population at the 2020 United States Census was 14,195, up from 11,491 at the 2010 census. It is part of the Jacksonville metropolitan stat ...
, the Wildlife Safari from
Winston, Oregon Winston is a city in Douglas County, Oregon, United States, located 7 miles southwest of Roseburg. The population was 5,379 according to the 2010 United States Census. Winston is best known as the home of Wildlife Safari. History In 1920, ...
, that bred more than 178 cheetahs and the De Wildt Cheetah and Wildlife Centre from South Africa where hundreds of cheetah cubs have been born. The Fota Wildlife Park from
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
is also known for successfully breeding cheetahs in captivity right before starting a captive-breeding project with the
Northeast African cheetah The Northeast African cheetah (''Acinonyx jubatus soemmeringii'') is a cheetah subspecies occurring in Northeast Africa. Contemporary records are known in South Sudan, Uganda, and Ethiopia, but population status in Eritrea, Djibouti, Somalia, Ke ...
.


Gallery

File:Cheetah at Sunset.jpg, A cheetah silhouetted against a fiery sunset in Okavango Delta,
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
File:TheCheethcat.jpg, At
Sabi Sand Game Reserve Sabi Sand Game Reserve consists of a group of private game reserves located adjacent to the Kruger National Park in the Lowveld of Mpumalanga, South Africa. Its name comes from the Sabie River on its southern boundary and the Sand River flow ...
File:Gepard Portrait.jpg, At Farm ''Achalm'' in
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
File:Naankuse 3 cheetah release (7-9) 094b.jpg, At Naankuse Wildlife Sanctuary


In popular culture

* The book '' How It Was with Dooms'' tells the true story of a family raising an orphaned East African cheetah cub named Duma (the Swahili word for cheetah) in Kenya. The films ''Cheetah'' (1989) and ''Duma'' (2005) were both loosely based on this book. However, ''Duma'' takes place in South Africa instead of Kenya. The cheetahs that starred in the film were South African cheetahs from the Kragga Kamma Game Park of the
Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...
province. In November 2011, one of the five adult cheetahs that starred in the film had died from an unusual kidney failure. * The
Toyota is a Japanese Multinational corporation, multinational Automotive industry, automotive manufacturer headquartered in Toyota City, Aichi, Japan. It was founded by Kiichiro Toyoda and incorporated on August 28, 1937. Toyota is the List of manuf ...
Free State Cheetahs The Free State Cheetahs (), formerly known as Orange Free State (), currently named the Toyota Free State Cheetahs due to sponsorship reasons, are a South African rugby union team that participates in the annual Currie Cup tournament. They are ...
, founded in 1895, is a South African
rugby union Rugby union football, commonly known simply as rugby union in English-speaking countries and rugby 15/XV in non-English-speaking world, Anglophone Europe, or often just rugby, is a Contact sport#Terminology, close-contact team sport that orig ...
team that participates in the annual
Currie Cup The Currie Cup () is South Africa's premier domestic rugby union competition featuring teams representing either entire provinces or substantial regions within provinces. Although it is the premier domestic competition, four South African franc ...
tournament. They have a cheetah running at high speed as their emblem. * The
Cheetahs The cheetah (''Acinonyx jubatus'') is a large cat and the fastest land animal. It has a tawny to creamy white or pale buff fur that is marked with evenly spaced, solid black spots. The head is small and rounded, with a short snout and blac ...
are another South African rugby union team from
Bloemfontein Bloemfontein ( ; ), also known as Bloem, is the capital and the largest city of the Free State (province), Free State province in South Africa. It is often, and has been traditionally, referred to as the country's "judicial capital", alongsi ...
founded in 2005 that have a running cheetah as their emblem.


See also

* Asiatic cheetah * Northwest African cheetah *
Northeast African cheetah The Northeast African cheetah (''Acinonyx jubatus soemmeringii'') is a cheetah subspecies occurring in Northeast Africa. Contemporary records are known in South Sudan, Uganda, and Ethiopia, but population status in Eritrea, Djibouti, Somalia, Ke ...
*
Big cat The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. All cats descend from the ''Felidae'' family, sharing similar musculature, c ...


References


External links

* Species portrai
''Acinonyx jubatus''
IUCN/SSC Cat Specialist Group
Cheetah Conservation Fund

Southern cheetah (''Acinonyx jubatus jubatus'')
{{Taxonbar, from=Q20721337
southeast African cheetah The Southeast African cheetah (''Acinonyx jubatus jubatus'') is the nominate subspecies, nominate cheetah subspecies native to East Africa, East and Southern Africa. The Southern African cheetah lives mainly in the Kalahari Basin, lowland areas a ...
Mammals of East Africa Mammals of Southern Africa Mammals of Angola Mammals of Botswana Mammals of Kenya Mammals of Malawi Mammals of Mozambique Mammals of Namibia Mammals of Somalia Mammals of South Africa Mammals of Eswatini Mammals of Tanzania Mammals of Uganda Mammals of Zambia Mammals of Zimbabwe
southeast African cheetah The Southeast African cheetah (''Acinonyx jubatus jubatus'') is the nominate subspecies, nominate cheetah subspecies native to East Africa, East and Southern Africa. The Southern African cheetah lives mainly in the Kalahari Basin, lowland areas a ...