Sorraia
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The Sorraia is a rare
breed A breed is a specific group of breedable domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist seve ...
of
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
indigenous to the portion of the
Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...
, in the Sorraia River basin, in
Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
. The Sorraia is known for its primitive features, including a convex profile and
dun Dun most commonly refers to: *Dun gene, which produces a brownish-gray color (dun) in horses and other Equidae * Dun (fortification), an ancient or medieval fort Dun or DUN may also refer to: Places Scotland * Dun, Angus, a civil parish in ...
coloring with
primitive markings Primitive markings are a group of equine coat color, hair coat markings and qualities seen in several equine species, including horses, donkey, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such ...
. Concerning its origins, a theory has been advanced by some authors that the Sorraia is a descendant of primitive horses belonging to the naturally occurring wild
fauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...
of Southern
Iberia The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, compri ...
. Studies are currently ongoing to discover the relationship between the Sorraia and various wild horse types, as well as its relationship with other breeds from the Iberian Peninsula and Northern Africa. Members of the breed are small, but hardy and well-adapted to harsh conditions. They were captured and used by native farmers for centuries, and a remnant population of these nearly
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
horses was discovered by a Portuguese
zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...
in the early 20th century. Today, the Sorraia has become the focus of preservation efforts, with European scientists leading the way and enthusiasts from several countries forming projects and establishing herds to assist in the re-establishment of this breed from its current endangered status.


Characteristics

The Sorraia breed stands between high, although some individuals are as small as The head tends to be large, the profile convex, and the ears long. The neck is slender and long, the
withers Withers are the ridge between the shoulder blades of an animal, typically a quadruped. In many species, this ridge is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, catt ...
high, and the
croup Croup ( ), also known as croupy cough, is a type of respiratory infection that is usually caused by a virus. The infection leads to swelling inside the trachea, which interferes with normal breathing and produces the classic symptoms of "bar ...
slightly sloping. The legs are strong, with long
pastern The pastern is a part of the leg of a horse between the fetlock and the top of the hoof. It incorporates the Equine_forelimb_anatomy#Metacarpal_bones, long pastern bone (proximal phalanx) and the Equine_forelimb_anatomy#Middle_phalanx, short past ...
s and well-proportioned hooves. These horses have good endurance and are
easy keeper An easy keeper, easy doer, or (British English) good doer is a horse that can live on relatively little feed. The opposite of an easy keeper is a '' hard keeper'' (''poor doer''), an animal that is prone to be too thin and has difficulty mainta ...
s, thriving on relatively little
fodder Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food ...
. They have a reputation for being independent of temperament, but tractable. On adult horses, the lay of the hair can create the appearance of stars and flags on the neck and chest. Also due to the lay of the hair, newborn
foal A foal is an equine up to one year old; this term is used mainly for horses, but can be used for donkeys. More specific terms are colt (horse), colt for a male foal and filly for a female foal, and are used until the horse is three or four. Whe ...
s can appear to have stripes all over, reminiscent of
zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: Grévy's zebra (''Equus grevyi''), the plains zebra (''E. quagga''), and the mountain zebra (''E. ...
stripes. The breed standard refers to this as "hair stroke".


Color

Sorraia are generally
dun Dun most commonly refers to: *Dun gene, which produces a brownish-gray color (dun) in horses and other Equidae * Dun (fortification), an ancient or medieval fort Dun or DUN may also refer to: Places Scotland * Dun, Angus, a civil parish in ...
or a dun variation called
grullo Grullo (pronounced GREW-yo) or grulla is a color of horses in the dun family, characterized by tan-gray or mouse-colored hairs on the body, often with shoulder and dorsal stripes and black barring on the lower legs. The genotype for grulla horse ...
. Dun coloring includes
primitive markings Primitive markings are a group of equine coat color, hair coat markings and qualities seen in several equine species, including horses, donkey, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such ...
such as a black dorsal stripe, black tipped ears, horizontal striping on the legs and a dark muzzle area. The dark muzzle area is in contrast to some other dun-colored horse breeds, who have light-colored muzzle areas and underbellies, possibly due to the presence of pangare genetics. Sorraia horses have bi-colored manes and tails with lighter colored hairs that fringe the outside of the longer growing black hair. This is a characteristic shared with other predominantly dun-colored breeds, such as the
Fjord horse The Fjord or Norwegian Fjord Horse () is a relatively small but very strong horse breed from the mountainous regions of western Norway. It is an agile breed of light draught horse build. It is always dun in colour, with five variations in sha ...
. Purebred Sorraia occasionally have
white markings Markings on horses are usually distinctive white areas on an otherwise dark base coat color. Most horses have some markings, and they help to identify the horse as a unique individual. Markings are present at birth and do not change over the ...
, although they are rare and undesired by the breed's
studbook A breed registry, also known as a herdbook, studbook or register, in animal husbandry, the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeders w ...
.


Genetics

The relationship between the Sorraia and other
breeds A breed is a specific group of breedable domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist seve ...
remains largely undetermined, as is its relationship to the
wild horse The wild horse (''Equus ferus'') is a species of the genus Equus (genus), ''Equus'', which includes as subspecies the modern domestication of the horse, domesticated horse (''Equus ferus caballus'') as well as the Endangered species, endangered ...
subspecies, the
Tarpan The tarpan (''Equus ferus ferus'') was a free-ranging horse population of the Eurasian steppe from the 18th to the 20th century. What qualifies as a tarpan is subject to debate; it is unclear whether tarpans were genuine wild horses, feral domest ...
and the
Przewalski's Horse Przewalski's horse (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered wild horse originally native to the steppes of Central Asia. It is named after t ...
. The Sorraia originally developed in the southern part of the Iberian Peninsula. d'Andrade hypothesized that the Sorraia would be the ancestor of the Southern Iberian breeds. Morphologically, scientists place the Sorraia as closely related to the Gallego and the Asturcon, but genetic studies using
mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
show that the Sorraia forms a cluster that is largely separated from most Iberian breeds. Some evidence links this cluster with
Konik The Konik or Polish Konik, , is a Polish list of horse breeds, breed of small horse or pony. There are semi-feral populations in some regions. They are usually dun gene, mouse dun or primitive markings, striped dun. The Bilgoray, , of south-e ...
and domestic Mongolian horses. At the same time, one of the maternal lineages is shared with the
Lusitano The Lusitano, also known as the Pure Blood Lusitano or PSL (''Puro Sangue Lusitano''), is a Portuguese horse breed. Horses were known to be present on the Iberian Peninsula as far back as 20,000 BC, and by 800 BC the region was renowned for ...
. Genetic evidence has not supported a hypothesis that the Sorraia is related to the
Barb horse The Barb horse, also known as the Barbary horse (), is a breed of riding horse with historic roots in North Africa. Known for its hardiness and stamina, it has influenced a number of modern breeds, including many in northern and western Afri ...
, an African breed introduced to Iberia by the Moors. Multiple authors have suggested that the Sorraia might be a descendant of the
Tarpan The tarpan (''Equus ferus ferus'') was a free-ranging horse population of the Eurasian steppe from the 18th to the 20th century. What qualifies as a tarpan is subject to debate; it is unclear whether tarpans were genuine wild horses, feral domest ...
based on shared morphological features, principally the typical color of its coat. Other authors simply state that the Sorraia has "evident primitive characteristics", although they do not refer to a specific ancestor. However, there have been no genetic studies comparing the Sorraia with the Tarpan, and similarity of external morphology is an unreliable measure of relatedness. Genetic studies to date have been inconclusive about the closest relative of the Sorraia. On one hand, studies using mitochondrial DNA showed a relationship with the
Przewalski's Horse Przewalski's horse (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered wild horse originally native to the steppes of Central Asia. It is named after t ...
, in that Przewalski's Horse has a unique haplotype (A2) not found in domestic horses, which differs by just one single
nucleotide Nucleotides are Organic compound, organic molecules composed of a nitrogenous base, a pentose sugar and a phosphate. They serve as monomeric units of the nucleic acid polymers – deoxyribonucleic acid (DNA) and ribonucleic acid (RNA), both o ...
from one of the major Sorraia haplotypes (JSO41, later A7). In comparison, genetic distances within the domestic horse are as large as 11 nucleotide differences. However, this relationship with the Przewalski's Horse was contradicted in another study using
microsatellite A microsatellite is a tract of repetitive DNA in which certain Sequence motif, DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organ ...
data that showed that the genetic distance between the Prewalski's Horse and the Sorraia was the largest. Such conflicting results can arise when a population passes through a
genetic bottleneck A population bottleneck or genetic bottleneck is a sharp reduction in the size of a population due to environmental events such as famines, earthquakes, floods, fires, disease, and droughts; or human activities such as genocide, speciocide, wid ...
, and evidence suggests that the Sorraia, among other rare breeds, has recently passed through a bottleneck, effectively obscuring the position of this breed in the family tree of the domestic horse. Thus, the morphological,
physiological Physiology (; ) is the science, scientific study of function (biology), functions and mechanism (biology), mechanisms in a life, living system. As a branches of science, subdiscipline of biology, physiology focuses on how organisms, organ syst ...
, and cultural characteristics of the Sorraia are the subject of continued study to better understand the relationship between various Iberian horse breeds and wild horse subspecies.


History

Although it is known that the Sorraia developed in the southern part of the Iberian Peninsula, the breed was isolated and unknown to science until the 20th century. Despite the lack of documentation, attempts have been made to reconstruct its history.
Paleolithic The Paleolithic or Palaeolithic ( years ago) ( ), also called the Old Stone Age (), is a period in human prehistory that is distinguished by the original development of stone tools, and which represents almost the entire period of human prehist ...
parietal art In archaeology, rock arts are human-made markings placed on natural surfaces, typically vertical stone surfaces. A high proportion of surviving historic and prehistoric rock art is found in caves or partly enclosed rock shelters; this type al ...
images in the region depict equines with a distinct likeness to the Sorraia, with similar zebra-like markings. Analysis of
mtDNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...
has been performed on
Mustang The mustang is a free-roaming horse of the Western United States, descended from horses brought to the Americas by the Spanish conquistadors. Mustangs are often referred to as wild horses, but because they are descended from once-domesticate ...
s in the western United States that show similar mtDNA patterns between some Mustangs and the Sorraia breed. Spanish
conquistador Conquistadors (, ) or conquistadores (; ; ) were Spanish Empire, Spanish and Portuguese Empire, Portuguese colonizers who explored, traded with and colonized parts of the Americas, Africa, Oceania and Asia during the Age of Discovery. Sailing ...
s took
Iberian horse The Iberian horse is a designation given to a number of horse breeds native to the Iberian Peninsula. At present, some breeds are officially recognized by the FAO,
s, some of whom closely resembled the modern-day Sorraia, to the Americas in their conquests, probably as pack animals. Similarities between the Sorraia and several North and South American breeds are shown in the dun and grullo coloring and various physical characteristics. This evidence suggests that the Sorraia, their ancestors, or other horses with similar features, may have had a long history in the Iberian region and a role in the creation of American breeds. Otherwise, the Sorraia breed was lost to history until 1920, when Portuguese zoologist and paleontologist Dr. Ruy d'Andrade first encountered the Sorraia horse during a hunting trip in the Portuguese lowlands. This remnant herd of primitive horses had continued to live a wild existence in these lowlands, which were rather inaccessible and had been used as a hunting preserve by Portuguese royalty until the early 1900s. At the time of d'Andrade's initial meeting the breed, the horses were ill-regarded by native farmers, although they were considered hardy native fauna that lived off of the uncultivated lands and salt marshes in the local river valleys. For centuries, peasant farmers of the area would occasionally capture the horses and use them for
agricultural Agriculture encompasses crop and livestock production, aquaculture, and forestry for food and non-food products. Agriculture was a key factor in the rise of sedentary human civilization, whereby farming of domesticated species created f ...
work, including
threshing Threshing or thrashing is the process of loosening the edible part of grain (or other crop) from the straw to which it is attached. It is the step in grain preparation after reaping. Threshing does not remove the bran from the grain. History of ...
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
and
herding Herding is the act of bringing individual animals together into a group (herd), maintaining the group, and moving the group from place to place—or any combination of those. Herding can refer either to the process of animals forming herds in ...
bull A bull is an intact (i.e., not Castration, castrated) adult male of the species ''Bos taurus'' (cattle). More muscular and aggressive than the females of the same species (i.e. cows proper), bulls have long been an important symbol cattle in r ...
s. In the 1920s and 1930s, as mechanization became more prevalent, both wild and domesticated breeding stock diminished to almost nothing, and d'Andrade, along with his son Fernando, encouraged the conservation of the breed. In 1937, d'Andrade began a small herd of his own with five
stallion A stallion is an adult male horse that has not been gelded ( castrated). Stallions follow the conformation and phenotype of their breed, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cre ...
s and seven
mare A mare is an adult female horse or other equidae, equine. In most cases, a mare is a female horse over the age of three, and a filly is a female horse three and younger. In Thoroughbred horse racing, a mare is defined as a female horse more th ...
s from horses obtained near
Coruche Coruche () is a municipality in Santarém District in Portugal. The population in 2011 was 19,944, in an area of 1115.72 km². The present Mayor is Francisco Silvestre de Oliveira, elected by the Socialist Party. The Coruche City Council ha ...
, Portugal. All Sorraias currently in captivity descend from these original horses obtained by d'Andrade, and it is believed that the remnant wild herds of the breed died out soon after. These horses were kept in a habitat similar to their native one. In 1975, two other farms took up the Sorraia's cause and acquired small herds to help with conservation. In 1976, three stallions and three mares were imported to Germany from Portugal to begin a sub-population there. In March 2004, a small breeding herd of Sorraia horses was released on the estate of a private land owner who dedicated a portion of his property so that these horses could live completely wild, as did their ancestors. The refuge created for them is in the Vale de Zebro region of south western Portugal, one of places so named because this is where the Sorraia's predecessors dwelt. Today, the breed is nearly extinct, with fewer than 200 horses existing as of 2007, including around 80 breeding mares. The
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations; . (FAO) is a specialized agency of the United Nations that leads international efforts to defeat hunger and improve nutrition and food security. Its Latin motto, , translates ...
considers it to be maintaining critical risk status. The first
studbook A breed registry, also known as a herdbook, studbook or register, in animal husbandry, the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeders w ...
was published in 2004, dedicated to maintaining a written record of the bloodlines of the Sorraia. Sorraias are present mainly in Portugal, with a small population in Germany. While not bred for a specific use, the Sorraia horses are versatile and have been used in herding bulls,
dressage Dressage ( or ; , most commonly translated as "training") is a form of horse riding performed in exhibition and competition, as well as an art sometimes pursued solely for the sake of mastery. As an equestrianism, equestrian sport defined by th ...
riding and light harness.


American preservation efforts

Two Sorraia stallions were imported to the United States in the early 21st century. In 2006, another Sorraia stallion was imported to Canada where a Sorraia Mustang Preserve has been established on
Manitoulin Island Manitoulin Island ( ) is an island in Lake Huron, located within the borders of the Provinces and territories of Canada, Canadian province of Ontario, in the bioregion known as Laurentia (bioregion), Laurentia. With an area of , it is the Lake ...
in Ontario. Unrelated to existing preservation efforts which work in conjunction with the Sorraia Mustang Studbook, another project by a consortium of breeders in the United States is attempting to establish a separate network and studbook. These breeders have gathered
Spanish Mustang The Spanish Mustang is an American horse breed descended from horses brought from Spain during the early conquest of the Americas. They are classified within the larger grouping of the Colonial Spanish horse, a type that today is rare in Spain. ...
s that through
mtDNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...
testing show a genetic relationship with the Sorraia and are breeding them according to both
genotype The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
and
phenotype In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology (physical form and structure), its developmental processes, its biochemical and physiological propert ...
in an attempt to help preserve what they are calling the "American Sorraia Mustang".


Naming

Dr. Ruy d'Andrade gave the breed their name of "Sorraia". D'Andrade took the name from the Sorraia River in Portugal. The breed had previously been known by the local Portuguese as "zebro" or "zebra", due to their markings. In the time of
Christopher Columbus Christopher Columbus (; between 25 August and 31 October 1451 – 20 May 1506) was an Italians, Italian explorer and navigator from the Republic of Genoa who completed Voyages of Christopher Columbus, four Spanish-based voyages across the At ...
, the Sorraia was also known as the Marismeño, but the Sorraia and the Marismeño have evolved into two different breeds over time. Today, the name Marismeño refers to a population of semiferal horses living in Doñana Natural Park in Spain.


References


External links


A page dedicated to the Sorraia.
{{good article Horse breeds Feral horses Horse breeds originating in Portugal