The Sonoran Desert tortoise (''Gopherus morafkai''), or Morafka's desert tortoise, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of terrestrial

chelonian Turtles are reptiles of the order Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), ...

reptile of the family

Testudinidae Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like ot ...

(tortoises) native to the

Sonoran Desert The Sonoran Desert () is a hot desert and ecoregion in North America that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the Southwestern United States (in Arizona and California). It ...

Taxon history

In 2011, using DNA evidence, geography, and behavioral observations, it was concluded that two distinct species of desert tortoise exist on either side (east-west) of the

Colorado River The Colorado River () is one of the principal rivers (along with the Rio Grande) in the Southwestern United States and in northern Mexico. The river, the List of longest rivers of the United States (by main stem), 5th longest in the United St ...

: *the Mojave or Agassiz's desert tortoise (''Gopherus agassizii''); known primarily from the

California counties The U.S. state of California is divided into 58 counties. The state was first divided into 27 counties on February 18, 1850. These were further sub-divided to form sixteen additional counties by 1860. Another fourteen counties were formed throu ...

of Imperial, Inyo,

Kern Kern or KERN may refer to: People * Kern (surname), includes a list of people with the name * Kern (soldier), a light infantry unit in Medieval Irish armies Places * Kern, Alaska, a ghost town in Alaska * Kern, Austria, see Sankt Marienkir ...

Los Angeles Los Angeles, often referred to by its initials L.A., is the List of municipalities in California, most populous city in the U.S. state of California, and the commercial, Financial District, Los Angeles, financial, and Culture of Los Angeles, ...

, Riverside,

San Bernardino San Bernardino ( ) is a city in and the county seat of San Bernardino County, California, United States. Located in the Inland Empire region of Southern California, the city had a population of 222,101 in the 2020 census, making it the List of ...

, and

San Diego San Diego ( , ) is a city on the Pacific coast of Southern California, adjacent to the Mexico–United States border. With a population of over 1.4 million, it is the List of United States cities by population, eighth-most populous city in t ...

. Its range continues northeast of

Death Valley Death Valley is a desert valley in Eastern California, in the northern Mojave Desert, bordering the Great Basin Desert. It is thought to be the Highest temperature recorded on Earth, hottest place on Earth during summer. Death Valley's Badwat ...

and the

Mojave Desert The Mojave Desert (; ; ) is a desert in the rain shadow of the southern Sierra Nevada mountains and Transverse Ranges in the Southwestern United States. Named for the Indigenous peoples of the Americas, indigenous Mohave people, it is located pr ...

through the

Nevada counties There are 16 counties and one independent city in the U.S. state of Nevada. On November 25, 1861, the first Nevada Territorial Legislature established nine counties. Nevada was admitted to the Union on October 31, 1864, with 11 counties. In 196 ...

Clark Clark is an English language surname with historical links to England, Scotland, and Ireland, ultimately derived from the Latin ''clericus'' meaning "scribe", "secretary" or a scholar within a religious order, referring to someone who was educated ...

, Esmeralda,

Lincoln Lincoln most commonly refers to: * Abraham Lincoln (1809–1865), the 16th president of the United States * Lincoln, England, cathedral city and county town of Lincolnshire, England * Lincoln, Nebraska, the capital of Nebraska, U.S. * Lincoln (na ...

and Nye, as well as

Mohave County Mohave County occupies the northwestern corner of the U.S. state of Arizona, one of 15 counties in the state. As of the 2020 census, its population was 213,267. The county seat is Kingman, and the largest city is Lake Havasu City. It is th ...

, in

Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...

, and extreme southwestern

Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...

, near

Zion National Park Zion National Park is a national park of the United States located in southwestern Utah near the town of Springdale. Located at the junction of the Colorado Plateau, Great Basin, and Mojave Desert regions, the park has a unique geography and a ...

. *the Sonoran or Morafka's desert tortoise (''Gopherus morafkai'');Genetic Analysis Splits Desert Tortoise into Two Species
US Geological Survey (2011-06-28). Retrieved on 2019-08-22. found east of the Colorado River, primarily in the Arizona counties of

Cochise Cochise ( ; Apache: or , ; later or , ; June 8, 1874) was the leader of the Chiricahui local group of the Chokonen and principal nantan of the Chokonen band of a Chiricahua Apache. A key war leader during the Apache Wars, he led an upri ...

, Gila,

Graham Graham or Graeme may refer to: People * Graham (given name), an English-language given name * Graham (surname), an English-language surname * Graeme (surname), an English-language surname * Graham (musician) (born 1979), Burmese singer * Clan ...

La Paz La Paz, officially Nuestra Señora de La Paz (Aymara language, Aymara: Chuqi Yapu ), is the seat of government of the Bolivia, Plurinational State of Bolivia. With 755,732 residents as of 2024, La Paz is the List of Bolivian cities by populati ...

Maricopa Maricopa can refer to: Places * Maricopa, Arizona, United States, a city in Pinal County, Arizona ** Maricopa Freeway, a section of I-10 in Metropolitan Phoenix ** Maricopa station, an Amtrak station in Maricopa, Arizona * Maricopa County, Arizo ...

, Mohave, Pima, Pinal, Santa Cruz,

Yavapai The Yavapai ( ) are a Native American tribe in Arizona. Their Yavapai language belongs to the Upland Yuman branch of the proposed Hokan language family. Today Yavapai people are enrolled in the following federally recognized tribes: * Fort ...

and Yuma. Found in the

, in the states of

Sonora Sonora (), officially Estado Libre y Soberano de Sonora (), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is divided into Municipalities of Sonora, 72 ...

and

Sinaloa, Mexico Sinaloa (), officially the (), is one of the 31 states which, along with Mexico City, compose the Federal Entities of Mexico. It is divided into 18 municipalities, and its capital city is Culiacán Rosales. It is located in northwest Mexic ...

. The new species' name is in honor of the late Professor David Joseph Morafka of

California State University, Dominguez Hills California State University, Dominguez Hills (CSUDH, CSU Dominguez Hills, or Cal State Dominguez Hills) is a public university in Carson, California. It was founded in 1960 and is part of the California State University (CSU) system. In 2020, ...

, in recognition of his many contributions to the study and conservation of the genus ''Gopherus''.

References

{{Taxonbar , from=Q2113453 Gopherus Turtles of North America Reptiles of Mexico Reptiles of the United States Reptiles described in 2011 Taxa named by Amy Lathrop Taxa named by Robert W. Murphy Taxa named by Alan E. Leviton