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The Solnhofen Limestone or Solnhofen Plattenkalk, formally known as the Altmühltal Formation, is a
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
Konservat-Lagerstätte that preserves a rare assemblage of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
ized organisms, including highly detailed imprints of soft bodied organisms such as sea jellies. The most familiar fossils of the Solnhofen Plattenkalk include the early feathered
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur ''
Archaeopteryx ''Archaeopteryx'' (; ), sometimes referred to by its German name, "" ( ''Primeval Bird'') is a genus of bird-like dinosaurs. The name derives from the ancient Greek (''archaîos''), meaning "ancient", and (''ptéryx''), meaning "feather" ...
'' preserved in such detail that they are among the most famous and most beautiful fossils in the world. The Solnhofen beds lie in the German state of Bavaria (Bayern), halfway between Nuremberg (Nürnberg) and Munich (München) and were originally quarried as a source of
lithographic limestone Lithographic limestone is hard limestone that is sufficiently fine-grained, homogeneous and defect-free to be used for lithography. Geologists use the term "lithographic texture" to refer to a grain size under 1/250 mm. The term "sublitho ...
. The Jura Museum situated in
Eichstätt Eichstätt () is a town in the federal state of Bavaria, Germany, and capital of the district of Eichstätt. It is located on the Altmühl river and has a population of around 13,000. Eichstätt is also the seat of the Roman Catholic Dioce ...
, Germany has an extensive exhibit of
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
fossils from the quarries of Solnhofen and surroundings, including marine reptiles,
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
s, and one specimen of the early bird ''
Archaeopteryx ''Archaeopteryx'' (; ), sometimes referred to by its German name, "" ( ''Primeval Bird'') is a genus of bird-like dinosaurs. The name derives from the ancient Greek (''archaîos''), meaning "ancient", and (''ptéryx''), meaning "feather" ...
''.


Paleoenvironment and preservation

During the
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
, this area was an
archipelago An archipelago ( ), sometimes called an island group or island chain, is a chain, cluster, or collection of islands. An archipelago may be in an ocean, a sea, or a smaller body of water. Example archipelagos include the Aegean Islands (the o ...
at the edge of the Tethys Sea. This included placid lagoons that had limited access to the open sea and where salinity rose high enough that the resulting brine could not support life. Since the lowest water was devoid of
oxygen Oxygen is a chemical element; it has chemical symbol, symbol O and atomic number 8. It is a member of the chalcogen group (periodic table), group in the periodic table, a highly reactivity (chemistry), reactive nonmetal (chemistry), non ...
, many ordinary scavengers were absent. Any organism that fell, drifted, or was washed into the lagoons from the ocean or the land became buried in soft
carbonate A carbonate is a salt of carbonic acid, (), characterized by the presence of the carbonate ion, a polyatomic ion with the formula . The word "carbonate" may also refer to a carbonate ester, an organic compound containing the carbonate group ...
mud. Thus, many delicate creatures avoided consumption by scavengers or being torn apart by currents. The wings of
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
, the imprints of stray
feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and an exa ...
s, and terrestrial plants that washed into the lagoons were all preserved. The fossils are not numerous, but some of them are spectacular, and their range gives a comprehensive picture of a local Jurassic
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
. At times, the lagoons almost dried out, exposing sticky carbonate muds that trapped insects and a few small
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s. Over 600 species have been identified, including twenty-nine kinds of
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
ranging from the size of a sparrow to 1.2 m (4 ft) in length. The fine-grained texture of the mud
silt Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension (chemistry), suspension with water. Silt usually ...
forming the
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
from the
Solnhofen Solnhofen is a municipality in the district of Weißenburg-Gunzenhausen in the region of Middle Franconia in the ' of Bavaria in Germany. It is in the Altmühl valley. The local area is famous in geology and palaeontology for Solnhofen lime ...
area (which is composed mainly of the towns of
Solnhofen Solnhofen is a municipality in the district of Weißenburg-Gunzenhausen in the region of Middle Franconia in the ' of Bavaria in Germany. It is in the Altmühl valley. The local area is famous in geology and palaeontology for Solnhofen lime ...
and
Eichstätt Eichstätt () is a town in the federal state of Bavaria, Germany, and capital of the district of Eichstätt. It is located on the Altmühl river and has a population of around 13,000. Eichstätt is also the seat of the Roman Catholic Dioce ...
) is ideal for making lithographic plates, and extensive quarrying in the 19th century revealed many fossil finds, as commemorated in the name ''
Archaeopteryx lithographica ''Archaeopteryx'' (; ), sometimes referred to by its German name, "" ( ''Primeval Bird'') is a genus of bird-like dinosaurs. The name derives from the ancient Greek (''archaîos''), meaning "ancient", and (''ptéryx''), meaning "feather" ...
'', all the specimens of which come from these deposits. The closest modern analogue to the Solnhofen conditions is said to be Orca Basin in the northern
Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...
, though that area is much deeper than the Solnhofen lagoons.


IUGS geological heritage site

In respect of the locality being a 'first-class paleontological site displaying high-quality reference material for scientists worldwide, including all known specimens of ''Archaeopteryx'' ', the
International Union of Geological Sciences The International Union of Geological Sciences (IUGS) is an international non-governmental organization devoted to global cooperation in the field of geology. As of 2023, it represents more than 1 million geoscientists around the world. About Fo ...
(IUGS) included the 'Jurassic Solnhofen-Eichstätt Archaeopteryx Serial Site' in its assemblage of 100 'geological heritage sites' around the world in a listing published in October 2022. The organisation defines an IUGS Geological Heritage Site as 'a key place with geological elements and/or processes of international scientific relevance, used as a reference, and/or with a substantial contribution to the development of geological sciences through history.'


Paleobiota

Significant members of the Solnhofen paleofauna include ''
Archaeopteryx ''Archaeopteryx'' (; ), sometimes referred to by its German name, "" ( ''Primeval Bird'') is a genus of bird-like dinosaurs. The name derives from the ancient Greek (''archaîos''), meaning "ancient", and (''ptéryx''), meaning "feather" ...
'', pterosaurs, and marine invertebrates.


Psilomelane dendrites

Dendrite A dendrite (from Ancient Greek language, Greek δένδρον ''déndron'', "tree") or dendron is a branched cytoplasmic process that extends from a nerve cell that propagates the neurotransmission, electrochemical stimulation received from oth ...
s, branched dark brown to black
fractal In mathematics, a fractal is a Shape, geometric shape containing detailed structure at arbitrarily small scales, usually having a fractal dimension strictly exceeding the topological dimension. Many fractals appear similar at various scale ...
tree-structure, are quite common in the limestone. Such formations are not plant fossil remains, but
iron Iron is a chemical element; it has symbol Fe () and atomic number 26. It is a metal that belongs to the first transition series and group 8 of the periodic table. It is, by mass, the most common element on Earth, forming much of Earth's o ...
and manganese oxides (
psilomelane Psilomelane is a group name for hard black manganese oxides including hollandite and romanechite. Psilomelane consists of hydrous manganese oxide with variable amounts of barium and potassium. Psilomelane is erroneously, and uncommonly, known as ...
, ) that crystallized out of solutions on bedding planes of the Solnhofen limestone. Dendrites have a chemical-mineralogical origin and are formed by a
diffusion Diffusion is the net movement of anything (for example, atoms, ions, molecules, energy) generally from a region of higher concentration to a region of lower concentration. Diffusion is driven by a gradient in Gibbs free energy or chemical p ...
process.


Cultural items made from Solnhofen Limestone

File:1514 Krug Adam und Eva Sündenfall The Fall of Man Bodemuseum anagoria.JPG, Adam and Eve (
Fall of Man The fall of man, the fall of Adam, or simply the Fall, is a term used in Christianity to describe the transition of the first man and woman from a state of innocent obedience to God in Christianity, God to a state of guilty disobedience. * * * * ...
), 1514 by Ludwig Krug, Bode-Museum
Berlin Berlin ( ; ) is the Capital of Germany, capital and largest city of Germany, by both area and List of cities in Germany by population, population. With 3.7 million inhabitants, it has the List of cities in the European Union by population withi ...
File:1525 Hering Liebesgarten - Jungbrunnen anagoria.JPG, ''Garden of Love/Fountain of Youth'', 1525 by Loy Hering ( Bode-Museum Berlin) File:1530 Kayser Susanna und die beiden Alten anagoria.JPG, ''Susannah and the Elders'', 1530 by Victor Kayser ( Bode-Museum Berlin) File:1648 Schweigger Salome with Head of St. John the Baptist anagoria.JPG, Salome Receives the Head of St. John the Baptist in the Dungeon, 1648 by Georg Schweigger


See also

* Jura Museum * List of fossil sites ''(with link directory)'' * List of types of limestone *
List of dinosaur-bearing rock formations This list of dinosaur-bearing rock formations is a list of geologic formations in which dinosaur fossils have been documented. * List of stratigraphic units with dinosaur body fossils * List of stratigraphic units with dinosaur trace fossils ** ...
* South German Jurassic


References


External links

{{Commons category, Solnhofen limestone
University of California Berkeley offers a brief introduction


Jurassic System of Europe Limestone formations Lagoonal deposits Fossiliferous stratigraphic units of Europe Paleontology in Germany Weißenburg-Gunzenhausen First 100 IUGS Geological Heritage Sites