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The snow petrel (''Pagodroma nivea'') is the only member of the genus ''Pagodroma.'' It is one of only three birds that have been seen at the Geographic
South Pole The South Pole, also known as the Geographic South Pole or Terrestrial South Pole, is the point in the Southern Hemisphere where the Earth's rotation, Earth's axis of rotation meets its surface. It is called the True South Pole to distinguish ...
, along with the Antarctic petrel and the south polar skua, which has the most southerly breeding sites of any bird, inland in Antarctica.


Taxonomy

The snow petrel was described in 1777 by the German naturalist
Georg Forster Johann George Adam Forster, also known as Georg Forster (; 27 November 1754 – 10 January 1794), was a German geography, geographer, natural history, naturalist, ethnology, ethnologist, travel literature, travel writer, journalist and revol ...
in his book ''
A Voyage Round the World ''A Voyage Round the World'' (complete title ''A Voyage Round the World in His Britannic Majesty's Sloop, Resolution, Commanded by Capt. James Cook, During the Years 1772, 3, 4, and 5'') is Georg Forster's report on the second voyage of the B ...
''. He had accompanied
James Cook Captain (Royal Navy), Captain James Cook (7 November 1728 – 14 February 1779) was a British Royal Navy officer, explorer, and cartographer famous for his three voyages of exploration to the Pacific and Southern Oceans, conducted between 176 ...
on Cook's second voyage to the Pacific.
We particularly observed a petrel, about the size of a pigeon, entirely white, with a black bill and blueish feet; it constantly appeared about the icy masses, and may be looked upon as a sure fore-runner of ice.
Forster placed the snow petrel in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Procellaria Procellaria is a genus of Southern Ocean long-winged seabirds related to prions, and within the order Procellariiformes. The black petrel (''Procellaria parkinsoni'') ranges in the Pacific Ocean, and as far north as Central America. The spec ...
'' that had been erected for the petrels by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1758 and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Procellaria nivea''. The snow petrel is now the only species placed in the genus ''Pagodroma'' that was introduced for the snow petrel in 1856 by French naturalist
Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857) was a French naturalist and ornithology, ornithologist, and a nephew of Napoleon. Lucien and his wife had twelve children, including Cardinal ...
. The genus name combines the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''pagos'' meaning "frost" or "sea-ice" with ''dromos'' meaning "racer" or "runner". The specific epithet is from the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''niveus'' meaning "snow-white". The word "petrel" is derived from
Peter the Apostle Saint Peter (born Shimon Bar Yonah; 1 BC – AD 64/68), also known as Peter the Apostle, Simon Peter, Simeon, Simon, or Cephas, was one of the Twelve Apostles of Jesus and one of the first leaders of the early Christian Church. He appears repe ...
and the story of his walking on water. This is in reference to the petrel's habit of appearing to run on the water to take off. Snow petrels vary significantly in size with two different forms, a larger and a smaller. The two forms hybridise extensively and this has led to uncertainty about the taxonomic status and the precise geographic distribution of the different types. In 1857 Bonaparte in his ''Conspectus Generum Avium'' listed subspecies ''major'' and ''minor'' but as he provided no further information, these names are not recognised and are considered as ''
nomen nudum In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. The German ornithologist
Hermann Schlegel Hermann Schlegel (10 June 1804 – 17 January 1884) was a German ornithologist, herpetologist and ichthyologist. Early life and education Schlegel was born at Altenburg, the son of a brassfounder. His father collected butterflies, which stimulated ...
in 1863 provided descriptions for the subspecies ''minor'' and ''major'' but erroneously believed that Forster's original description applied to the larger form. In 1912
Gregory Mathews Gregory Macalister Mathews CBE FRSE FZS FLS (10 September 1876 – 27 March 1949) was an Australian-born amateur ornithologist who spent most of his later life in England. Life He was born in Biamble in New South Wales the son of Robert H. M ...
, in the second volume of his ''Birds of Australia'', treated the larger form as a separate species and introduced the binomial name ''Pagodroma confusa''. The two forms are now usually treated as subspecies and Schlege is acknowledged as the authority for the larger subspecies. The snow petrel is a member of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Procellariidae The family (biology), family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prion (bird), prions, and the shearwaters. This family is part of the bird order (biology), orde ...
, and the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Procellariiformes Procellariiformes is an order (biology), order of seabirds that comprises four family (biology), families: the albatrosses, the Procellariidae, petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still call ...
. They all share certain identifying features. First, they have nasal passages that attach to the upper bill called
naricorns A nostril (or naris , : nares ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbinates, ...
. The bills of Procellariiformes are also unique in that they are split into seven to nine horny plates. They produce a
stomach oil Stomach oil is the light oil composed of neutral dietary lipids found in the proventriculus (fore-gut) of birds in the order Procellariiformes. All albatrosses, procellarids (gadfly petrels and shearwaters) and northern and austral storm ...
made up of
wax ester A wax ester (WE) is an ester of a fatty acid and a fatty alcohol. Wax esters are the main components of three commercially important waxes: carnauba wax, candelilla wax, and beeswax.. Wax esters are formed by combining one fatty acid with one ...
s and
triglycerides A triglyceride (from ''wikt:tri-#Prefix, tri-'' and ''glyceride''; also TG, triacylglycerol, TAG, or triacylglyceride) is an ester derived from glycerol and three fatty acids. Triglycerides are the main constituents of body fat in humans and oth ...
that is stored in the
proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects. Birds Th ...
. This can be sprayed out of their mouths as a defense against predators (principally
skua The skuas are a group of predatory seabirds with seven species forming the genus ''Stercorarius'', the only genus in the family Stercorariidae. The three smaller skuas, the Arctic skua, the long-tailed skua, and the pomarine skua, are called ...
s) and as an energy-rich food source for the chicks and for the adults during their long flights. Finally, they also have a
salt gland The salt gland is an organ (anatomy), organ for excreting excess salt (chemistry), salts. It is found in the cartilaginous fishes subclass elasmobranchii (sharks, rays, and skates), seabirds, and some reptiles. Salt glands can be found in the r ...
situated above the nasal passage which helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a high saline solution from their nostrils.


Subspecies

Two subspecies are recognised: * ''P. n. nivea'' ( Forster, G, 1777) — breeds on the
Antarctic Peninsula The Antarctic Peninsula, known as O'Higgins Land in Chile and Tierra de San Martin in Argentina, and originally as Graham Land in the United Kingdom and the Palmer Peninsula in the United States, is the northernmost part of mainland Antarctica. ...
,
South Georgia Islands South Georgia and the South Sandwich Islands (SGSSI) is a British Overseas Territory in the southern Atlantic Ocean. It is a remote and inhospitable collection of islands, consisting of South Georgia and a chain of smaller islands known as the S ...
, and other islands of the Scotia Arc * ''P. n. major'' ( Schlegel, 1863), formerly ''P. n. confusa'' ( Mathews, 1912) — breeds on the
South Sandwich Islands The South Sandwich Islands () are a chain of uninhabited volcanic islands in the South Atlantic Ocean. They are administered as part of the British Overseas Territory of South Georgia and the South Sandwich Islands. The chain lies in the sub-A ...
and
Géologie Archipelago The Géologie Archipelago, also known as the Pointe Géologie Archipelago, Geology Archipelago or Cape Geology Archipelago, is a small archipelago of rocky islands and rocks close to the north of Cape Géodésie and Astrolabe Glacier Tongue, ext ...


Description

The snow petrel is a small, pure white fulmarine petrel with coal-black eyes, a small black bill and bluish gray feet. Body length is and the wingspan is . Flight is erratic with frequent changes of direction.


Behavior

Snow petrels are almost entirely restricted to cold Antarctic waters. Flocks are characteristically seen sitting on icebergs.


Breeding

Breeding occurs in
colonies A colony is a territory subject to a form of foreign rule, which rules the territory and its indigenous peoples separated from the foreign rulers, the colonizer, and their '' metropole'' (or "mother country"). This separated rule was often or ...
on the Antarctic continent and on various Antarctic islands. Nesting is colonial in small to large colonies on exposed rocks, usually near the sea, but also on inland mountain ranges more than from the open sea. Some birds remain at the colony all year, but the main influx at colonies is from the mid-September until early November. Nests are simple pebble-lined scrapes usually in a deep rock crevices with overhanging protection. Nests are abandoned if concealed by heavy snowfall; egg mortality is 50%, and chick mortality is 10–15%. One white
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
is laid between late November and mid-December. The egg is incubated 41 to 49 days and the chick is brooded for 8 days. They
fledge Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
7 weeks later in late February to mid-May. Colonies are also the sites of cleaning areas where snow petrels, far from the sea, bathe in snow. Snow petrel partners are faithful for life (around 20 years).


Feeding

Snow petrels feed mainly on fish, some
cephalopod A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...
s,
mollusks Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The num ...
, and
krill Krill ''(Euphausiids)'' (: krill) are small and exclusively marine crustaceans of the order (biology), order Euphausiacea, found in all of the world's oceans. The name "krill" comes from the Norwegian language, Norwegian word ', meaning "small ...
, as well as
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
in the form of seal
placenta The placenta (: placentas or placentae) is a temporary embryonic and later fetal organ that begins developing from the blastocyst shortly after implantation. It plays critical roles in facilitating nutrient, gas, and waste exchange between ...
s, dead/stillborn seals, whale carcasses, and dead penguin chicks. During the winter, they disperse to the
pack ice Pack or packs may refer to: Music * Packs (band), a Canadian indie rock band * ''Packs'' (album), by Your Old Droog * ''Packs'', a Berner album Places * Pack, Styria, defunct Austrian municipality * Pack, Missouri, United States (US) * ...
, ice floes, and the open sea. Flocks are characteristically seen sitting on icebergs. Only very rarely are they observed north of the pack ice.


Conservation

The snow petrel has an occurrence range of , and an estimated population of 4 million adult birds. Based on these numbers, the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has classified this bird as
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
.


References

{{Authority control
snow petrel The snow petrel (''Pagodroma nivea'') is the only member of the genus ''Pagodroma.'' It is one of only three birds that have been seen at the Geographic South Pole, along with the Antarctic petrel and the south polar skua, which has the most s ...
snow petrel The snow petrel (''Pagodroma nivea'') is the only member of the genus ''Pagodroma.'' It is one of only three birds that have been seen at the Geographic South Pole, along with the Antarctic petrel and the south polar skua, which has the most s ...
Birds of Antarctica Birds of islands of the Atlantic Ocean Birds of subantarctic islands
snow petrel The snow petrel (''Pagodroma nivea'') is the only member of the genus ''Pagodroma.'' It is one of only three birds that have been seen at the Geographic South Pole, along with the Antarctic petrel and the south polar skua, which has the most s ...
South Pole