
The Pyralidae, commonly called pyralid moths,
snout moths or grass moths, are a
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
Lepidoptera in the
ditrysia
The Ditrysia are a natural group or clade of insects in the lepidopteran order containing both butterflies and moths. They are so named because the female has two distinct sexual openings: one for mating, and the other for laying eggs (in cont ...
n
superfamily
SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Pyraloidea
The Pyraloidea (pyraloid moths or snout moths) are a moth superfamily containing about 16,000 described species worldwide, and probably at least as many more remain to be described. They are generally fairly small moths, and as such, they have b ...
. In many (particularly older) classifications, the
grass moth
The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies inclu ...
s (Crambidae) are included in the Pyralidae as a
subfamily, making the combined group one of the largest families in the Lepidoptera. The latest review by
Eugene G. Munroe and
Maria Alma Solis
Maria Alma Solis is a entomologist at the Systematic Entomology Laboratory (SEL) of the Agricultural Research Service of the U.S. Department of Agriculture.
Early life
Maria Alma Solis was born on February 9, 1956, in Corpus Christi, Texas, ...
retain the
Crambidae
The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies includ ...
as a full family of
Pyraloidea
The Pyraloidea (pyraloid moths or snout moths) are a moth superfamily containing about 16,000 described species worldwide, and probably at least as many more remain to be described. They are generally fairly small moths, and as such, they have b ...
.
The
wingspan
The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan ...
s for small and medium-sized species are usually between with variable morphological features.
It is a diverse group, with more than 6,000 species described worldwide, and more than 600 species in America north of Mexico, comprising the third largest moth family in North America. At least 42 species have been recorded from North Dakota in the subfamilies of Pyralidae.
Relationship with humans
Most of these small moths are inconspicuous. Many are economically important pests, including
waxworm
Waxworms are the caterpillar larvae of wax moths, which belong to the family Pyralidae (snout moths). Two closely related species are commercially bred – the lesser wax moth (''Achroia grisella'') and the greater wax moth (''Galleria ...
s, which are the
caterpillar
Caterpillars ( ) are the larva, larval stage of members of the order Lepidoptera (the insect order comprising butterfly, butterflies and moths).
As with most common names, the application of the word is arbitrary, since the larvae of sawfly ...
larva
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
Th ...
e of the greater (''
Galleria mellonella
''Galleria mellonella'', the greater wax moth or honeycomb moth, is a moth of the family Pyralidae. ''G. mellonella'' is found throughout the world. It is one of two species of wax moths, with the other being the lesser wax moth. ''G. mellonella ...
'') and lesser (''
Achroia grisella
The lesser wax moth (''Achroia grisella'') is a small moth of the snout moth family (Pyralidae) that belongs to the subfamily Galleriinae. The species was first described by Johan Christian Fabricius in 1794. Adults are about 0.5 inches (13&nbs ...
'') wax moths (
subfamily Galleriinae
The Galleriinae are a subfamily of snout moths (family Pyralidae) and occur essentially worldwide, in some cases aided by involuntary introduction by humans. This subfamily includes the wax moths, whose caterpillars (waxworms) are bred on a com ...
). They are natively
pest
Pest or The Pest may refer to:
Science and medicine
* Pest (organism), an animal or plant deemed to be detrimental to humans or human concerns
** Weed, a plant considered undesirable
* Infectious disease, an illness resulting from an infection
** ...
s of
beehive
A beehive is an enclosed structure in which some honey bee species of the subgenus ''Apis'' live and raise their young. Though the word ''beehive'' is commonly used to describe the nest of any bee colony, scientific and professional literature ...
s, but are bred indoors in enormous numbers as
live food
Live food is living animals used as food for other carnivorous or omnivorous animals kept in captivity; in other words, small preys (such as insects, small fish or rodents) fed alive to larger predators kept either in a zoo or as a pet.
Live ...
for small
reptile and
bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
pet
A pet, or companion animal, is an animal kept primarily for a person's company or entertainment rather than as a working animal, livestock, or a laboratory animal. Popular pets are often considered to have attractive appearances, intelligence ...
s and similar animals. They are also used as
fishing bait
Fishing bait is any substance used to attract and catch fish, e.g. on a fishing hook. Bait items are both selected from and placed within the environment to achieve enhanced prey capture success. Traditionally, fishing baits are natural fish foo ...
for
trout
Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', '' Salmo'' and '' Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-sa ...
fishing.
Other notable snout moth pests relevant for their larval hosts include:
* Alligatorweed stem borer (''
Arcola malloi
''Arcola malloi'' (formerly ''Vogtia malloi'') is a species of Pyralidae, snout moth known as the alligator weed stem borer. It is used as an agent of biological pest control against the noxious weed, noxious aquatic plant known as alligator weed ...
'':
Phycitinae
The Phycitinae are a subfamily of snout moths (family Pyralidae). Even though the Pyralidae subfamilies are all quite diverse, Phycitinae stand out even by standards of their family: with over 600 genera considered valid and more than 4000 spec ...
) –
biological control
Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically als ...
of alligator weed (''
Alternanthera philoxeroides'').
* Almond moth (''
Cadra cautella'': Phycitinae) – pest of stored
cereal
A cereal is any grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, germ, and bran. Cereal grain crops are grown in greater quantities and provide more foo ...
s and dry fruit; now introduced almost worldwide.
* Cacao moth, tobacco moth, warehouse moth (''
Ephestia elutella
''Ephestia elutella'', the cacao moth, tobacco moth or warehouse moth, is a small moth of the family Pyralidae. It is probably native to Europe, but has been transported widely, even to Australia. A subspecies is ''E. e. pterogrisella''.
The w ...
'': Phycitinae) – pest of stored dry vegetable products; Europe, introduced to some other regions (e.g. Australia).
* Dried fruit moth (''
Cadra calidella
''Cadra calidella'', the dried fruit or date moth, is a species of snout moth in the genus '' Cadra'' and commonly mistaken for the species '' Cadra figulilella.'' It thrives in warmer conditions and is found primarily in Mediterranean countries ...
'':
Phycitinae
The Phycitinae are a subfamily of snout moths (family Pyralidae). Even though the Pyralidae subfamilies are all quite diverse, Phycitinae stand out even by standards of their family: with over 600 genera considered valid and more than 4000 spec ...
)
* ''
Etiella behrii'' (Phycitinae) – pest of stored
legume
A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock fo ...
s; Southeast Asia and Australia
* "
Flour moths" – pests of stored grains, spices, flour, and similar dry vegetable products; now introduced almost worldwide.
** Indian mealmoth (''
Plodia interpunctella
The Indianmeal moth (''Plodia interpunctella''), also spelled Indian meal moth and Indian-meal moth, is a pyraloid moth of the family Pyralidae. Alternative common names are weevil moth, pantry moth, flour moth or grain moth. The almond m ...
'': Phycitinae)
** Mediterranean flour moth, Indian flour moth (''
Ephestia kuehniella
The Mediterranean flour moth or mill moth (''Ephestia kuehniella'') is a moth of the family Pyralidae. It is a common pest of cereal grains, especially flour. This moth is found throughout the world, especially in countries with temperate climate ...
'': Phycitinae)
* Grease moth (''
Aglossa pinguinalis
''Aglossa pinguinalis'', the large tabby or grease moth, is a moth in the subfamily Pyralinae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''.
The forewings are greyish brown clouded with a da ...
'':
Pyralinae
The Pyralinae are the typical subfamily of snout moths ( family Pyralidae) and occur essentially worldwide, in some cases aided by involuntary introduction by humans. They are rather rare in the Americas however, and their diversity in the Austr ...
) – pest of
suet
Suet is the raw, hard fat of beef, lamb or mutton found around the loins and kidneys.
Suet has a melting point of between 45 °C and 50 °C (113 °F and 122 °F) and congelation between 37 °C and 40 °C (98.6&n ...
and other oily food.
* Lesser cornstalk borer (''
Elasmopalpus lignosellus'': Phycitinae) – stalk pest of
corn
Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn ( North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. ...
(''Zea mays''); tropical and subtropical Americas, introduced to the Hawaiian Islands.
* Locust bean moth (''
Ectomyelois ceratoniae
''Ectomyelois ceratoniae'', the locust bean moth, more ambiguously known as " carob moth", is a moth of the family Pyralidae. It has a nearly cosmopolitan distribution.
Adults have forewings with a pale brown pattern, and plain white hindwings. ...
'': Phycitinae)
* Mahogany webworm (''
Macalla thyrsisalis
''Macalla thyrsisalis'', the mahogany webworm moth, is a species of snout moth in the genus ''Macalla''. It was described by Francis Walker in 1859. It is found in southern Florida, the Bahamas, Hispaniola, southern Mexico, Central America, Trin ...
'':
Epipaschiinae
The Epipaschiinae are a subfamily of snout moths (family Pyralidae). More than 720 species are known today, which are found mainly in the tropics and subtropics. Some occur in temperate regions, but the subfamily is apparently completely absent ...
) – defoliator pest of
mahogany trees (''Swietenia'');
Neotropics
The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone.
Definition
In bi ...
.
* Meal moth (''
Pyralis farinalis
''Pyralis'' is a genus of snout moths. It was described by Carl Linnaeus in 1758.
Species
* '' Pyralis caustica'' (Meyrick, 1884)
* '' Pyralis costinotalis'' Hampson, 1917
* '' Pyralis electalis'' Hulst, 1886
* '' Pyralis farinalis'' (Linnaeus, ...
'': Pyralinae) – pest of stored grain, flour and other cereals; now introduced almost worldwide.
* Pear fruit borer (''
Pempelia heringii'': Phycitinae) – pest of apple and pear fruits; East Asia, introduced to the Hawaiian Islands.
* Pine webworm (''
Pococera robustella'': Epipaschiinae) – defoliator pest of
pine
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanic ...
s (''Pinus''); North America east of Great Lakes region.
* Raisin moth (''
Cadra figulilella
''Cadra figulilella'', the raisin moth, is a moth of the family Pyralidae. The raisin moth is known most commonly as a pest that feeds on dried fruits, such as the raisin and date. It covers a range that includes much of the world, primarily situ ...
'': Phycitinae) – pests of stored dry fruit; now introduced almost worldwide.
* Rice moth (''
Corcyra cephalonica
The rice moth (''Corcyra cephalonica'') is a moth of the family Pyralidae. This small moth can become a significant pest. Its caterpillars feed on dry plantstuffs such as seeds, including cereals (e.g. rice). Other recordedGrabe (1942) foods ar ...
'': Galleriinae) – pest of stored grain, flour and other cereals.
* South American cactus moth (''
Cactoblastis cactorum
''Cactoblastis cactorum'', the cactus moth, South American cactus moth or nopal moth, is native to Argentina, Paraguay, Uruguay and southern Brazil. It is one of five species in the genus ''Cactoblastis'' that inhabit South America, where many par ...
'': Phycitinae) – biological control of
prickly pears (''Opuntia'').
* Southern pine coneworm, "pitch moth" (''
Dioryctria amatella
''Dioryctria amatella'', the southern pineconeworm moth, is a species of moth of the family Pyralidae. It is found in the south-eastern United States, from Maryland south to Florida and west into Texas.
The wingspan is 27–32 mm. The forew ...
'': Phycitinae) – cone and shoot pest of pines (''Pinus''); southern North America.
* Stored nut moth (''
Paralipsa gularis'': Galleriinae) – pest of stored nuts and
drupe
In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part ( exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kerne ...
s; Southeast Asia, introduced to Western Europe.
* Sunflower moth (''
Homoeosoma nebulella'': Phycitinae) – pest of
sunflower seed
The sunflower seed is the seed of the sunflower (''Helianthus annuus''). There are three types of commonly used sunflower seeds: linoleic (most common), high oleic, and sunflower oil seeds. Each variety has its own unique levels of monounsatu ...
s; Europe and surrounding regions.
The European corn borer (''
Ostrinia nubilalis'') and southern cornstalk borer (''
Diatraea crambidoides
''Diatraea crambidoides'', the southern cornstalk borer moth, is a species of moth of the family Crambidae described by Augustus Radcliffe Grote in 1880. It is found in North America, from Alabama and northern Florida to Ohio and Maryland. Its win ...
''), formerly considered snout moths, are placed in the
Crambidae
The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies includ ...
which, as noted above, are usually regarded as a separate family today.
Systematics

Five subfamilies are generally recognized in the Pyralidae today. The
Acentropinae
Acentropinae is a fairly small subfamily of the lepidopteran family Crambidae, the crambid snout moths. Species of this subfamily are exclusively found in wetlands and aquatic habitats.
Systematics
In modern treatments, the former sub ...
(= Nymphulinae), occasionally still placed here, do indeed seem to belong in the Crambidae.
The snout moth subfamilies are, listed in the presumed
phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
sequence from the most primitive to the most advanced:
*
Chrysauginae
The Chrysauginae are a subfamily of snout moths (family Pyralidae). They are primarily Neotropical and include about 400 described species.
Description and ecology
The subfamily includes the sloth moths (genera '' Cryptoses'', ''Bradypodicola ...
(including Bradypodicolinae, Semniidae) – about 400 species occurring predominantly in the
Neotropical
The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone.
Definition
In bi ...
region. Larvae typically feed on plants, but some have more unusual feeding habits. The latter include for example some
myrmecophilous
Myrmecophily ( , ) is the term applied to positive interspecies associations between ants and a variety of other organisms, such as plants, other arthropods, and fungi. Myrmecophily refers to mutualistic associations with ants, though in its m ...
species, as well as a number of
sloth moth
A sloth moth is a coprophagous moth which has evolved to exclusively inhabit the fur of sloths and to use sloth dung as a substrate for the early stages of reproduction. Sloth moths include '' Bradypodicola hahneli'', ''Cryptoses choloepi'', ''Cr ...
s which are dependent on
sloth
Sloths are a group of Neotropical xenarthran mammals constituting the suborder Folivora, including the extant arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of movement, tree sloths spend most of their li ...
s for their entire life cycle. Most Chrysauginae larvae have a sclerotised ring around seta SD1 of the
metathorax
The metathorax is the posterior of the three segments in the thorax of an insect, and bears the third pair of legs. Its principal sclerites ( exoskeletal plates) are the metanotum (dorsal), the metasternum (ventral), and the metapleuron (lateral) ...
.
*
Galleriinae
The Galleriinae are a subfamily of snout moths (family Pyralidae) and occur essentially worldwide, in some cases aided by involuntary introduction by humans. This subfamily includes the wax moths, whose caterpillars (waxworms) are bred on a com ...
(including Macrothecinae) – about 300 species worldwide. The males of galleriine moths have a gnathos almost or completely reduced, the pupae have a prominent dorsal median ridge on the thorax and abdomen, and most larvae have a sclerotised ring around seta SD1 of the first abdominal segment.
*
Pyralinae
The Pyralinae are the typical subfamily of snout moths ( family Pyralidae) and occur essentially worldwide, in some cases aided by involuntary introduction by humans. They are rather rare in the Americas however, and their diversity in the Austr ...
(including Endotrichinae, Hypotiinae) – rather diverse in the
Old World
The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by th ...
; a lesser number of the roughly 900 species occurs elsewhere. The females of almost all Pyralinae except ''
Cardamyla'' and ''
Embryoglossa
''Embryoglossa'' is a genus of snout moths. It was described by Warren in 1896, and is known from Nigeria, Madagascar, and India.
Species
* '' Embryoglossa aethiopicalis'' Gaede, 1916
* '' Embryoglossa bipuncta'' Hampson, 1903
* '' Embryoglos ...
'' are recognizable by the very short
ductus bursae
In anatomy and physiology, a duct is a circumscribed channel leading from an exocrine gland or organ.
Types of ducts
Examples include:
Duct system
As ducts travel from the acinus which generates the fluid to the target, the ducts become larger ...
of their
genital
A sex organ (or reproductive organ) is any part of an animal or plant that is involved in sexual reproduction. The reproductive organs together constitute the reproductive system. In animals, the testis in the male, and the ovary in the female, ...
s.
*
Epipaschiinae
The Epipaschiinae are a subfamily of snout moths (family Pyralidae). More than 720 species are known today, which are found mainly in the tropics and subtropics. Some occur in temperate regions, but the subfamily is apparently completely absent ...
(including Pococerinae) – over 550 described species in the tropical and temperate regions (except Europe). Larvae are leaf rollers, leaf tiers, or leaf miners. Some species are minor pests of a few commercial crops. Epipaschiinae are generally hard to recognize, except in the case of adult males which have a few characteristic traits, such as the upturned and pointed third segment of the labial palps and usually a scaly projection from the antenna base. The larvae lack any stereotyped seta sclerotisations.
*
Phycitinae
The Phycitinae are a subfamily of snout moths (family Pyralidae). Even though the Pyralidae subfamilies are all quite diverse, Phycitinae stand out even by standards of their family: with over 600 genera considered valid and more than 4000 spec ...
(including Anerastiinae, Peoriinae) – probably the most difficult group of Pyraloidea in terms of identification and classification. They comprise more than 600 genera and about 4000 species found all over the world. The characteristic trait of the caterpillars is a
sclerotised
Sclerotin is a component of the cuticle of various Arthropoda, most familiarly insects. It is formed by cross-linking members of particular classes of protein molecules, a biochemical process called sclerotization, a form of tanning in which qu ...
area encircling the base of seta SD1 on the
mesothorax
The mesothorax is the middle of the three segments of the thorax of hexapods, and bears the second pair of legs. Its principal sclerites (exoskeletal plates) are the mesonotum (dorsal), the mesosternum (ventral), and the mesopleuron (lateral) on ...
, while the adult females have – like the males of Pyralidae in general do – a frenulum consisting of a single bristle which in turn is composed of multiple
acanthae.
Problematic genera
In addition to those assigned to the tribes above, several genera of (presumed) Pyralidae are not firmly placed in this arrangement, but are ''
incertae sedis
' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
''. Some may be very
basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
lineages which stand outside the main snout moth radiations, but given the changing circumscription of the Pyralidae, some are likely to be placed outside this group in its modern meaning, either in the Crambidae or in other lineages of basal
Obtectomera
The Obtectomera is a clade of macro-moths and butterflies, comprising over 100,000 species in at least 12 superfamilies.
Taxonomy
The Obtectomera includes the following 12 superfamilies:
* Whalleyanoidea Minet, 1991
* Thyridoidea Herrich-Sc ...
. Some may even belong to more ancient moth lineages, such as the
Alucitoidea
Aluctoidea is the superfamily of many-plumed and false plume moths. These small moths are most easily recognized by their wings. These each consist of many (typically more than 3) narrow strips of membrane around the major veins, instead of a con ...
or
Pterophoroidea. Finally, some of these (usually little-studied) genera possibly are
junior synonyms of genera described earlier. These genera are in the unranked category of the family Pyralidae.
The genera in question are:
* ''
Apocabimoides''
Neunzig & Goodson, 1992
* ''
Ardjuna
''Ardjuna'' is a genus of snout moths. It contains the species ''Ardjuna kresna''. It is found on northern Sumatra.
References
Pyralidae
Monotypic moth genera
Moths of Indonesia
Pyralidae genera
{{Pyralidae-stub ...
''
Roesler & Küppers, 1979
* ''
Cabimoides''
Neunzig & Goodson, 1992
* ''
Cryptophycita''
Roesler & Küppers, 1979
* ''
Cryptozophera''
Roesler & Küppers, 1979
* ''
Delcina''
Clarke, 1986 (Phycitinae?)
* ''
Eupassadena''
Neunzig & Goodson, 1992
* ''
Gomezmenoria''
Agenjo, 1966
* ''
Gunungodes''
Roesler & Küppers, 1981
* ''
Indocabnia''
Roesler & Küppers, 1981
* ''
Inverina''
Neunzig & Goodson, 1992
* ''
Kasyapa
Kashyapa ( sa, कश्यप}, ) is a revered Vedic sage of Hinduism., Quote: "Kasyapa (Rudra),(Vedic Seer)..." He is one of the Saptarishis, the seven ancient sages of the ''Rigveda''. Kashyapa is the most ancient and venerated rishi, al ...
''
Roesler & Küppers, 1981
* ''
Kaurava
''Kaurava'' is a Sanskrit term which refers to descendants of Kuru, a legendary king of India who is the ancestor of many of the characters of the epic ''Mahabharata''. Usually, the term is used for the 100 sons of King Dhritarashtra and his w ...
''
Roesler & Küppers, 1981
* ''
Kumbhakarna
Kumbhakarna (Sanskrit: कुम्भकर्ण, lit. ''pot-eared'') is a powerful rakshasa and younger brother of Ravana from the Hindu epic ''Ramayana''. Despite his gigantic size and appetite, he is described as a virtuous character a ...
''
Roesler & Küppers, 1981
* ''
Merangiria''
Roesler & Küppers, 1979
* ''
Ohigginsia''
Neunzig & Goodson, 1992
* ''
Pseudopassadena''
Neunzig & Goodson, 1992
* ''
Psorozophera''
Roesler & Küppers, 1979
These genera have been placed in the Pyralidae when these were still circumscribed ''
sensu lato
''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...
'' and are sometimes still treated thus, but actually they seem to belong in the Crambidae (see also ''
Micronix
''Micronix'' is a genus of moths of the family Crambidae. It contains only one species, ''Micronix nivalis'', which is found in Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivarian ...
'' and ''
Tanaobela
''Tanaobela'' is a genus of moths in the superfamily Pyraloidea containing only one species, ''Tanaobela chrysochlora'', which is found in Australia, where it has been recorded from Queensland. Its affiliations are disputed.
The wingspan
...
''):
* ''
Alphacrambus''
Bassi, 1995
* ''
Peniculimius''
Schoute, 1994
* ''
Steneromene''
Gaskin, 1986
* ''
Thopeutis''
Hübner, 1818
* ''
Yoshiyasua'' (formerly ''
Melanochroa
''Yoshiyasua'' is a grass moth genus (family Crambidae) of subfamily Musotiminae. Some authors have placed it in the snout moth family (Pyralidae), where all grass moths were once also included, but this seems to be in error. The genus contains o ...
''
Yohiyasu, 1985 ''nec'' Roeder, 1886: preoccupied
The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently.
* In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linn ...
)
References
External links
* Solis, M. Alma (2007)
"Phylogenetic studies and modern classification of the Pyraloidea (Lepidoptera)" ''Revista Colombiana de Entomología''. 33 (1): 1–8.
*
{{Taxonbar , from=Q248425
Pyralidae
Moth families