The smooth snake (''Coronella austriaca'')Street D (1979). ''The Reptiles of Northern and Central Europe''. London: B.T. Batsford Ltd. 268 pp. . is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of non-

venomous snake ''Venomous snakes'' are species of the suborder Serpentes that are capable of producing venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The venom is typically delivered by injection using hollow ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Colubridae Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. C ...

. The species is found in northern and central Europe, but also as far east as northern Iran. The

Reptile Database The Reptile Database is a scientific database that collects taxonomic information on all living reptile species (i.e. no fossil species such as dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared ...

recognizes two

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

as being valid, including the

nominotypical subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...

described here.

Description

Both sexes of ''C. austriaca'' grow to an average total length (including tail) of about to . Two specimens measuring have been recorded in Sweden, as well as one in Russia that was . The head has a

rostral scale The rostral scale, or rostral, in snakes and other scaled reptiles is the median plate on the tip of the snout that borders the mouth opening. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: ...

that is at least as deep as it is wide, creating a triangular indentation between the internasal scales (rarely separating them). The top of the head is covered with nine large plates. The

nasal scale In reptiles, the nasal scale refers to the scale that encloses the nostril. Mallow D, Ludwig D, Nilson G (2003). ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. . Somet ...

is often divided. There is one (rarely two) preoculars and two postoculars. The temporals number 2+2 or 2+3 (rarely 1+2). There are seven (rarely eight) upper labials, of which the third and fourth or fourth and fifth border the eye. In the middle part of the body there are 19 (rarely 17 or 21) rows of

dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...

. In contrast with many other snakes found in the region, these scales are flat (not keeled). This gives the snake a smooth texture to the touch, from which it gets its common name. The

ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...

number 150–164 in males and 162–200 in females. The anal scale is divided (rarely single) and the

subcaudal scales In snakes, the subcaudal scales are the enlarged plates on the underside of the tail. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Comstock Publishing Associates, a Division of Cornell University Press. (7t ...

are paired. Males have 54–70 subcaudal scales and females 40–76. The colour pattern consists of a brown, grey or reddish ground colourSteward JW (1971). ''The Snakes of Europe''. Cranbury, New Jersey: Associated University Press (Fairleigh Dickinson University Press). 238 pp. LCCCN 77-163307. . with two rows of small, rather indistinct dark spots running down the back towards the tail. In some cases, each pair of spots may be united toward the neck area, forming a series of cross-bars over the back. There is also a very indistinct series of dark spots running along each of the flanks. These four series of spots along the body overlay four parallel, rather shadowy stripes that also run down the back and flanks. On the top of the head is a dark marking which is often in the shape of a crown, giving rise to the generic name ''Coronella'' (which means

coronet In British heraldry, a coronet is a type of crown that is a mark of rank of non-reigning members of the royal family and peers. In other languages, this distinction is not made, and usually the same word for ''crown'' is used irrespective of ra ...

). A relatively thick dark stripe extends from each nostril, through the eye, and along the side of the head to a little beyond the neck. The upper labials are whitish, greyish-white or light brown, sometimes with darker spots. The tongue is reddish brown or dark red.

Biology

The smooth snake feeds on smaller animals, especially other reptiles. It subdues larger prey by constriction, although unlike true constrictors it does not kill by this method. Smooth snakes are

ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a "bridging" form of reproduction between egg-laying oviparity, oviparous and live-bearing viviparity, viviparous reproduction. Ovoviviparous animals possess embryos that develo ...

: the juveniles hatch out of eggs internally and are born live. In Britain it is restricted to

heathland A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and is characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a coole ...

habitats.

Geographic range

''Coronella austriaca'' is found from the

south of England Southern England, also known as the South of England or the South, is a sub-national part of England. Officially, it is made up of the southern, south-western and part of the eastern parts of England, consisting of the statistical regions of ...

through France and the

Low Countries The Low Countries (; ), historically also known as the Netherlands (), is a coastal lowland region in Northwestern Europe forming the lower Drainage basin, basin of the Rhine–Meuse–Scheldt delta and consisting today of the three modern "Bene ...

to northern Spain and Portugal, Germany, Norway and Sweden (as far north as latitude 63°), Latvia, Lithuania, Estonia, Switzerland, Austria, Italy and

Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...

(but not in

Corsica Corsica ( , , ; ; ) is an island in the Mediterranean Sea and one of the Regions of France, 18 regions of France. It is the List of islands in the Mediterranean#By area, fourth-largest island in the Mediterranean and lies southeast of the Metro ...

Sardinia Sardinia ( ; ; ) is the Mediterranean islands#By area, second-largest island in the Mediterranean Sea, after Sicily, and one of the Regions of Italy, twenty regions of Italy. It is located west of the Italian Peninsula, north of Tunisia an ...

), the western

Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...

and Greece, and

European Russia European Russia is the western and most populated part of the Russia, Russian Federation. It is geographically situated in Europe, as opposed to the country's sparsely populated and vastly larger eastern part, Siberia, which is situated in Asia ...

as far north as latitude 57°. In Asia, it is found from Turkey to

Azerbaijan Azerbaijan, officially the Republic of Azerbaijan, is a Boundaries between the continents, transcontinental and landlocked country at the boundary of West Asia and Eastern Europe. It is a part of the South Caucasus region and is bounded by ...

Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...

Armenia Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...

and northern Iran. It is absent in Denmark despite the species being found just south of the German border as well as in southern Sweden. The species is only known from Denmark from 6 specimens, all found between 1870 and 1914. In Finland, the species is found only in

Åland Åland ( , ; ) is an Federacy, autonomous and Demilitarized zone, demilitarised region of Finland. Receiving its autonomy by a 1920 decision of the League of Nations, it is the smallest region of Finland by both area () and population (30,54 ...

, and it is not common there.

Subspecies

References

Gallery

Image:Rézsikló (Coronella austriaca) feje.jpg, Head of ''C. austriaca''. Image:Rézsikló (Coronella austriaca)3.jpg, ''C. austriaca'' in

Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...

. File:Coronella austriaca s.jpg, ''C. austriaca'' in

Bosnia and Herzegovina Bosnia and Herzegovina, sometimes known as Bosnia-Herzegovina and informally as Bosnia, is a country in Southeast Europe. Situated on the Balkans, Balkan Peninsula, it borders Serbia to the east, Montenegro to the southeast, and Croatia to th ...

. Image:coronella_austriaca.jpg, ''C. austriaca'' curled up. Image:CoronellaAustriaca2.jpg, Subadult ''C. austriaca'' in northern

Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...

Description

Biology

Geographic range

Subspecies

See also

References

Further reading

Gallery

External links