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Sminthidae is a family of
mouse A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
-like jumping
rodents Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
. They are represented by only one extant
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
, ''
Sicista Birch mice (genus ''Sicista'') are small jumping rodents that resemble mice with long, tufted tails and very long hind legs, allowing for remarkable leaps. They are the only extant members of the family Sminthidae. They are native to Eurasian fo ...
'', represented by 19 species found throughout most of
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
, from
central Europe Central Europe is an area of Europe between Western Europe and Eastern Europe, based on a common historical, social and cultural identity. The Thirty Years' War (1618–1648) between Catholicism and Protestantism significantly shaped the ...
east to
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
, and south to southern China. However, they were much more diverse and had a much wider range in prehistoric times, having multiple genera and being found not only in Eurasia but also throughout North America, where they existed up to the early
Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
. They have a well-attested fossil record which dates as far back as the early
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
. They were formerly classified as the subfamily Sicistinae in the family
Dipodidae Jerboas (from ar, جربوع ') are hopping desert rodents found throughout North Africa and Asia, and are members of the family Dipodidae. They tend to live in hot deserts. When chased, jerboas can run at up to . Some species are preyed on ...
alongside the
jerboas Jerboas (from ar, جربوع ') are hopping desert rodents found throughout North Africa and Asia, and are members of the family Dipodidae. They tend to live in hot deserts. When chased, jerboas can run at up to . Some species are preyed on b ...
and jumping mice, but phylogenetic evidence supports all three of these belonging to distinct families, thus leaving only the jerboas in Dipodidae.


Extant species

* Genus ''
Sicista Birch mice (genus ''Sicista'') are small jumping rodents that resemble mice with long, tufted tails and very long hind legs, allowing for remarkable leaps. They are the only extant members of the family Sminthidae. They are native to Eurasian fo ...
'' ** Armenian birch mouse ''Sicista armenica'' **
Northern birch mouse The northern birch mouse (''Sicista betulina'') is a small rodent about 5 to 8 cm long (without the tail), weighing 5 to 13 g. It lives in northern Europe and Asia in forest and marsh zones. It hibernates in burrows. It eats shoots, grain ...
, ''Sicista betulina'' ** Caucasian birch mouse, ''Sicista caucasica'' **
Long-tailed birch mouse The long-tailed birch mouse (''Sicista caudata'') is a species of birch mouse found in Northeast Asia. It has been reported from the Ussuri region of Manchuria, Sakhalin Island and Primorsky Krai in Russia, and northern North Korea. It is liste ...
, ''Sicista caudata'' **
Tsimlyansk birch mouse Tsimlyansk (russian: Цимля́нск) is a town and the administrative center of Tsimlyansky District in Rostov Oblast, Russia, located on the right bank of the Don River, on the coast of Tsimlyansk Reservoir, northeast of Rostov-on-Don ...
, ''Sicista cimlanica'' ** Chinese birch mouse, ''Sicista concolor'' ** Kazbeg birch mouse, ''Sicista kazbegica'' ** Kluchor birch mouse, ''Sicista kluchorica'' **
Nordmann's birch mouse Nordmann's birch mouse (''Sicista loriger'') is a species of birch mouse in the family Sminthidae. It is named after Finnish biologist Alexander von Nordmann. It is native to eastern and southeastern Europe. Taxonomy It was long thought to ...
, ''Sicista loriger'' **
Altai birch mouse The Altai birch mouse (''Sicista napaea'') is a species of rodent in the family Sminthidae. It is native to Russia and Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central A ...
, ''Sicista napaea'' **
Gray birch mouse The gray birch mouse (''Sicista pseudonapaea'') is a species of rodent in the family Sminthidae. It is endemic to Kazakhstan, but possibly ranges into China. Its natural habitat is temperate forest A forest is an area of land dominated ...
, ''Sicista pseudonapaea'' **
Severtzov's birch mouse The Severtzov's birch mouse, or dark birch mouse (''Sicista severtzovi'') is a species of rodent in the family Sminthidae. It is endemic to East-European steppes (E Ukraine and S-W Russia Russia (, , ), or the Russian Federation, is a ...
, ''Sicista severtzovi'' ** Strand's birch mouse, ''Sicista strandi'' ** Southern birch mouse, ''Sicista subtilis'' **
Talgar birch mouse Talgar ( kk, Талғар, translit=Talğar, ; russian: Талгар) is a town in Almaty Region, southeastern Kazakhstan. It is the administrative center of Talgar District. The town is located between Almaty and Esik, 25 km from Almaty a ...
, ''Sicista talgarica'' **
Terskey birch mouse Teskey ( ky, Тескей, Teskey; zh, 泰尔斯凯 ) is a village in Batken Region of Kyrgyzstan. It is part of the Kadamjay District. Its population was 191 in 2021. Nearby towns and villages include Aydarken , image_shield = Co ...
, ''Sicista terskeica'' **
Tien Shan birch mouse The Tien Shan birch mouse (''Sicista tianshanica'') is a species of rodent in the family Sminthidae. It is found in China, Kazakhstan and Kyrgyzstan. Description The Tien Shan birch mouse grows to a length of about with a tail length of about ...
, ''Sicista tianshanica'' **
Hungarian birch mouse The Hungarian birch mouse (''Sicista trizona'') is a species of birch mouse in the family Sminthidae. It was once found throughout Central Europe and Romania, but is now only known from two isolated populations in Hungary and Transylvania in Rom ...
, ''Sicista trizona'' **
Zhetysu birch mouse Zhetysu, or Jeti-Suu ( kk, , Жетісу, pronounced ; ky, ''Jeti-Suu'', (), meaning "seven rivers"; also transcribed ''Zhetisu'', ''Jetisuw'', ''Jetysu'', ''Jeti-su'', ''Jity-su'', ''Жетысу'',, United States National Geospatial-I ...
, ''Sicista zhetysuica''


Extinct genera

These extinct genera are definitively known: * Genus †'' Allosminthus'' * Genus †'' Heosminthus'' * Genus †'' Heterosminthus'' * Genus †'' Gobiosminthus'' * Genus †'' Lophocricetus'' * Genus †'' Macrognathomys'' * Genus †'' Megasminthus'' * Genus †'' Miosicista'' * Genus †'' Parasminthus'' * Genus †'' Plesiosminthus'' * Genus †'' Schaubemys'' * Genus †'' Shamosminthus'' * Genus †'' Sinosminthus'' * Genus †'' Tyrannomys'' The
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
genera '' Primisminthus'' and '' Banyuesminthus'' could represent even older members of Sminthidae, although other studies speculate that they may be basal dipodoids.


References

{{Taxonbar, from=Q50319991 Sminthidae Rodent families Dipodoid rodents Extant Oligocene first appearances Taxa named by Johann Friedrich von Brandt