Small Mountain Ringlet
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The small mountain ringlet or mountain ringlet (''Erebia epiphron'') is a
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
of the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species ha ...
. It is found in mountainous regions of southern and central Europe.


Distribution

Mountain areas of
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).


Great Britain

It is Britain's only true
alpine Alpine may refer to any mountainous region. It may also refer to: Places Europe * Alps, a European mountain range ** Alpine states, which overlap with the European range Australia * Alpine, New South Wales, a Northern Village * Alpine National P ...
species of butterfly. The larva feeds on moor matgrass near
bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and musk ...
s and springs and the nectar-feeding adult visits
bilberry Bilberries () are Eurasian low-growing shrubs in the genus ''Vaccinium'' in the flowering plant family Ericaceae that bear edible, dark blue berries. They resemble but are distinct from North American blueberries. The species most often referre ...
,
tormentil ''Potentilla'' is a genus containing over 500 species of annual, biennial and perennial herbaceous flowering plants in the rose family, Rosaceae. Potentillas may also be called cinquefoils in English, but they have also been called five finge ...
and
heath bedstraw ''Galium saxatile'' or heath bedstraw is a species of flowering plant in the family Rubiaceae. It is related to cleavers. ''Galium saxatile'' is a perennial mat-forming herb, found on grassland, moors, heaths and woods. It can reach a height of ...
.


Life history

The pale cream eggs are laid singly, each female laying up to 70. The egg stage lasts two or three weeks. The larva is green with a double dorsal and a single lateral white or yellowish line. The third
instar An instar (, from the Latin '' īnstar'' 'form, likeness') is a developmental stage of arthropods, such as insects, which occurs between each moult (''ecdysis'') until sexual maturity is reached. Arthropods must shed the exoskeleton in order to ...
larvae hibernate in grass tussocks. They emerge in the spring and recommence feeding. Some larvae spend two years in this stage, the result of a late spring and short summer restricting growth. Recorded larval food plants are ''
Aira praecox ''Aira praecox'' is a species of grass known by several common names, including early hair-grass, yellow hairgrass and spike hairgrass. It is native to Europe, where it is found in dry, sandy places, on rocky outcrops, and in heath grassland. It ...
'' (Czechoslovakia, Spain), ''
Deschampsia cespitosa ''Deschampsia'' is a genus of plants in the grass family Poaceae, commonly known as hair grass or tussock grass. The genus is widespread across many countries.Festuca ovina ''Festuca ovina'', sheep's fescue or sheep fescue, is a species of grass. It is sometimes confused with hard fescue (''Festuca trachyphylla''). General description It is a perennial plant sometimes found in acidic ground, and in mountain pastur ...
'' (Spain), ''
Nardus stricta ''Nardus'' is a genus of plants belonging to the grass family, containing the single species ''Nardus stricta'', known as matgrass. It is placed in its own tribe Nardeae within the subfamily Pooideae. The name derives from ancient Greek ' () fro ...
'' (British Isles) and ''
Poa annua ''Poa annua'', or annual meadow grass (known in America more commonly as annual bluegrass or simply poa), is a widespread low-growing turfgrass in temperate climates. Notwithstanding the reference to annual plant in its name, perennial bio-types ...
'' (Spain) The pupa is formed deep in grass tussocks, encased within a loose silk structure. The pupa stage lasts about three weeks. The flight period is extremely short, a few weeks only, June to August depending on altitude.


Description

Description in South (1906).


Adult

Wingspan . "The typical form of this butterfly, ''epiphron'', Knock, has the tawny bands unbroken on the fore wings, and almost so on the hind wings; the black dots on the hind wings of the female are often pupilled with white, and more rarely this is so in the male also. It has been stated that specimens occur in Perthshire which exhibit these characters. All the British examples of the small mountain ringlet that I have seen are referable to the form known as ''cassiope'', Fab. (Plate 77). The tawny, or orange, bands are rarely so entire on the fore wings as in ''epiphron'', and are generally rather narrower; and that on the hind wing is broken up into three or four rings. The black dots are usually smaller and without white pupils. The female is somewhat larger and the bands or rings paler. Variation in the markings is extensive. The bands on the fore wings become less and less complete, until they are reduced to a series of mere rings around the black dots. The black dots decrease in size and in number until they, together with the tawny marking, entirely disappear, and a plain blackish-brown insect only remains. This extreme form has been named ''obsoleta'', Tutt. The earliest rings to vanish seem to be the third on the fore wings and the first on the hind wings. Similar modifications occur on the underside also, but there may be an aberration on the upper side of a specimen, and not, or at least not in the same way, on the underside." Wheeler (1903) gives a short description Wheeler, George.''The butterflies of Switzerland and the alps of central Europe''. London: Elliot Stock, 1903
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Early stages

"The egg when first laid, is yellow, changing afterwards to fawn color with darker markings, especially towards the top. It is laid in July on blades of grass. The larva hatches in about sixteen days. The young caterpillar, before hibernation in October, is greenish, with darker green and yellow lines. Head brownish. Feeds in July and after hibernation on various grasses, among which ''Poa annua'', ''Festuca ovina'', ''Aira praecox'', and ''A. caespitosa'' have been specified as eaten by caterpillars in confinement. A distinct preference, however, has been shown for mat grass (''Nardus strictus''), and it has been suggested that this may be the natural food. The full-grown caterpillar appears to be undescribed. The chrysalis is described by Buckler as being "little more than three-eighths of an inch in length, rather thick in proportion, being less dumpy in form than ''hyperanthtis'', but more so than ''blandina'' 'Erebia cethiops'' The color of the back of the thorax and wing cases is a light green, rather glaucous; the abdomen a pale drab or dirty whitish; a dark brown dorsal streak is conspicuous on the thorax, and there is the faintest possible indication of its being continued as a stripe along the abdomen. The eye-, trunk-, antenna-, and leg-cases are margined with dark brown, and the wing nervures are indicated by the same colors."


Subspecies

*''Erebia epiphron epiphron'' Harz Mountains *''Erebia epiphron aetheria'' Esper, 1805 *''Erebia epiphron mackeri'' Fuchs, 1914 (synonym ''Erebia epiphron vögesiaca'' Goltz, 1914) *''Erebia epiphron mnemon'' Haworth, 1812 (synonym ''Erebia epiphron scotica'' Cooke, 1943) *''Erebia epiphron orientpyreanica'' Eisner, 1946 (synonym ''Erebia epiphron fauveani'') Pyrénées-Orientales *''Erebia epiphron pyreanica'' Herrich-Schläffer, 1851 *''Erebia epiphron silasiana'' Meyer-Dür, 1852 *''Erebia epiphron transylvanica'' Rebel, 1908 *''Erebia epiphron retyezatensis'' Warren 1931 *''Erebia epiphron roosi'' Arnscheid & Sterba *''Erebia epiphron orientalis'' or '' Erebia orientalis'' Bulgaria *''Erebia epiphron nelamus'' high altitude Hybrid ''Erebia serotina'' ''epiphron'' × '' Erebia pronoe''


References


External links


Lepiforum
{{Taxonbar, from=Q1809089 Erebia Butterflies of Europe Butterflies described in 1783