The engrailed and small engrailed (''Ectropis crepuscularia'') are

moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...

s of the family

Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyleti ...

found from the

British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...

through central and eastern Europe to the

Russian Far East The Russian Far East ( rus, Дальний Восток России, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in North Asia. It is the easternmost part of Russia and the Asia, Asian continent, and is coextensive with the Far Easte ...

and

Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...

. The western

Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...

and

Asia Minor Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...

and the

Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...

represent the southern limit of the distribution (with the

Balkan countries The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...

). In the north, the distribution area ends at the Arctic Circle. It also occurs in North America. Debate exists as to whether they make up one

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

, or whether ''E. crepuscularia'' actually refers only to the small engrailed, with the engrailed proper being separable as ''E. bistortata''. The ground colour of the wings is buff or grey, variably marked with darker

fascia A fascia (; : fasciae or fascias; adjective fascial; ) is a generic term for macroscopic membranous bodily structures. Fasciae are classified as superficial, visceral or deep, and further designated according to their anatomical location. ...

and a pale postdiscal crossline. The darker markings are not usually as strong as in the rather similar

willow beauty The willow beauty (''Peribatodes rhomboidaria'') is a moth of the family Geometridae. It is a common species of Europe and adjacent regions (Near East and the Maghreb). While it is found widely throughout Scandinavian countries, which have a mari ...

. Melanic forms occur fairly frequently. The

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...

is . One or two broods are produced each year. In the British Isles, the adults can be seen at any time between March and August; this time range may vary in other parts of this moth's range. The species flies at night and is attracted to light. The greyish

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...

is truly

polyphagous Feeding is the process by which organisms, typically animals, obtain food. Terminology often uses either the suffixes -vore, -vory, or -vorous from Latin ''vorare'', meaning "to devour", or -phage, -phagy, or -phagous from Greek φαγε ...

, feeding on a huge range of

plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...

s. As a caterpillar, the species is known as the saddleback looper. The species overwinters as a

pupa A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...

Recorded food plants

Ectropis crepuscularia caterpillar - Keila

* ''Acer'',

maple ''Acer'' is a genus of trees and shrubs commonly known as maples. The genus is placed in the soapberry family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated si ...

* ''

Aconitum ''Aconitum'' (), also known as aconite, monkshood, wolfsbane, leopard's bane, devil's helmet, or blue rocket, is a genus of over 250 species of flowering plants belonging to the family (biology), family Ranunculaceae. These herbaceous perennial ...

'', monkshood * ''Alnus'',

alder Alders are trees of the genus ''Alnus'' in the birch family Betulaceae. The genus includes about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few species ex ...

* ''

Aquilegia ''Aquilegia'', commonly known as columbines, is a genus of perennial flowering plants in the family Ranunculaceae (buttercups). The genus includes between 80 and 400 taxa (described species and subspecies) with natural Species distribution, rang ...

'', columbine * ''Betula'',

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains 3 ...

* ''

Calluna vulgaris ''Calluna vulgaris'', common heather, ling, or simply heather, is the sole species in the genus ''Calluna'' in the flowering plant family Ericaceae. It is a low-growing evergreen shrub growing to tall, or rarely to and taller, and is found wide ...

'', heather * ''

Camellia japonica ''Camellia japonica'', known as common camellia, or Japanese camellia, is a species of Camellia, a flowering plant genus in the family Theaceae. There are thousands of cultivars of ''C. japonica'' in cultivation, with many colors and forms of flo ...

'', Japanese camellia * ''Castanea'',

chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. The name also refers to the edible nuts they produce. They are native to temperate regions of the Northern Hemisphere. Description ...

* ''Centaurea'', star thistle * ''

Cirsium arvense ''Cirsium arvense'' is a perennial species of flowering plant in the family Asteraceae, native throughout Europe and western Asia, northern Africa and widely introduced elsewhere.Joint Nature Conservation Committee''Cirsium arvense'' The standa ...

'', creeping thistle * ''Cornus'',

dogwood ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods or cornels, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous ...

* ''

Daphniphyllum ''Daphniphyllum'' is the sole genus in the flowering plant family Daphniphyllaceae and was described as a genus in 1826. The genus includes evergreen shrubs and trees mainly native to east and southeast Asia, but also found in the Indian Subconti ...

'' * ''

Diervilla ''Diervilla'', or bush honeysuckle, is a genus of three species of deciduous shrubs in the family Caprifoliaceae, all indigenous to eastern North America. The genus is named after a France, French surgeon Dr. Marin Diereville, who introduced the ...

'', bush honeysuckle * ''Frangula'',

alder buckthorn ''Frangula alnus'', commonly known as alder buckthorn, glossy buckthorn, or breaking buckthorn, is a tall deciduous shrub in the family Rhamnaceae. Unlike other "buckthorns", alder buckthorn does not have thorns. It is native to Europe, northern ...

* ''Fraxinus'', ash * ''Genista'',

broom A broom (also known as a broomstick) is a cleaning tool, consisting of usually stiff fibers (often made of materials such as plastic, hair, or corn husks) attached to, and roughly parallel to, a cylindrical handle, the broomstick. It is thus a ...

* ''

Glycine Glycine (symbol Gly or G; ) is an amino acid that has a single hydrogen atom as its side chain. It is the simplest stable amino acid. Glycine is one of the proteinogenic amino acids. It is encoded by all the codons starting with GG (G ...

'', soybean * '' Hypericum maculatum'', imperforate St. John's wort * ''Ilex'',

holly ''Ilex'' () or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen o ...

* ''Juglans'',

walnut A walnut is the edible seed of any tree of the genus '' Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. They are accessory fruit because the outer covering of the fruit is technically an i ...

* ''Larix'',

Larch Larches are deciduous conifers in the genus ''Larix'', of the family Pinaceae (subfamily Laricoideae). Growing from tall, they are native to the cooler regions of the northern hemisphere, where they are found in lowland forests in the high la ...

* ''

Lindera Dried fruits of ''Lindera neesiana'' used as spice (coll. MHNT) ''Lindera'' is a genus of about 80–100honeysuckle Honeysuckles are arching shrubs or Vine#Twining vines, twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae. The genus includes 158 species native to northern latitudes in North America, Eurasia, and North Africa. Widely kno ...

* ''Lythrum salicaria'',

purple loosestrife ''Lythrum salicaria'' or purple loosestrifeFlora of NW Europe''Lythrum salicaria'' is a flowering plant belonging to the family Lythraceae. It should not be confused with other plants sharing the name loosestrife that are members of the family Pr ...

* ''

Malus ''Malus'' ( or ) is a genus of about 32–57 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples (sometimes known in North America as crabapples) and wild apples. The genus i ...

'', apple * '' Philadelphus'', mock-orange * ''Picea'',

spruce A spruce is a tree of the genus ''Picea'' ( ), a genus of about 40 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal ecosystem, boreal (taiga) regions of the Northern hemisphere. ''Picea'' ...

* '' Pieris'' * ''Pinus'',

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...

* ''

Plectranthus ''Plectranthus'' is a genus of about 85 species of flowering plants from the sage family, Lamiaceae, found mostly in southern and tropical Africa and Madagascar. Common names include spur-flower. ''Plectranthus'' species are herbaceous perennial ...

'' * ''

Populus ''Populus'' is a genus of 25–30 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar (), aspen, and cottonwood. The we ...

'', poplar * ''Pseudotsuga'',

Douglas fir The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is the tallest tree in the Pinaceae family. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Or ...

* ''Quercus'',

oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...

* ''Ribes rubrum'',

redcurrant The redcurrant or red currant (''Ribes rubrum'') is a member of the genus ''Ribes'' in the gooseberry family. It is native to western Europe. The species is widely cultivated and has escaped into the wild in many regions. Description ''Ribes ...

* ''Rosa'',

rose A rose is either a woody perennial plant, perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred Rose species, species and Garden roses, tens of thousands of cultivar ...

* ''Rubus idaeus'',

raspberry The raspberry is the edible fruit of several plant species in the genus ''Rubus'' of the Rosaceae, rose family, most of which are in the subgenus ''Rubus#Modern classification, Idaeobatus''. The name also applies to these plants themselves. Ras ...

Hübner (1825) ''Ectropis''
/ref> * ''

Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distri ...

'', dock * ''Salix'',

willow Willows, also called sallows and osiers, of the genus ''Salix'', comprise around 350 species (plus numerous hybrids) of typically deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions. Most species are known ...

* ''Sambucus'', elder * ''Shepherdia canadensis'', Canada buffaloberry * ''Sorbus'' spp.,

rowan The rowans ( or ) or mountain-ashes are shrubs or trees in the genus ''Sorbus'' of the rose family, Rosaceae. They are native throughout the cool temperate regions of the Northern Hemisphere, with the highest species diversity in the Himalaya ...

s * ''Spartina'',

cordgrass ''Spartina'' is a genus of plants in the grass family, frequently found in coastal salt marshes. Species in this genus are commonly known as cordgrass or cord-grass, and are native to the coasts of the Atlantic Ocean in western and southern Eu ...

* ''

Thuja ''Thuja'' ( ) is a genus of coniferous tree or shrub in the Cupressaceae (cypress family). There are five species in the genus, two native to North America and three native to eastern Asia. The genus is monophyletic and sister to ''Thujopsis''. M ...

'' * ''Trifolium'',

clover Clovers, also called trefoils, are plants of the genus ''Trifolium'' (), consisting of about 300 species of flowering plants in the legume family Fabaceae originating in Europe. The genus has a cosmopolitan distribution with the highest diversit ...

* ''

Tsuga ''Tsuga'' (, from Japanese (), the name of '' Tsuga sieboldii'') is a genus of conifers in the subfamily Abietoideae of Pinaceae, the pine family. The English-language common name "hemlock" arose from a perceived similarity in the smell of it ...

'', hemlock * ''Ulmus'',

elm Elms are deciduous and semi-deciduous trees comprising the genus ''Ulmus'' in the family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical- montane regions of North America and Eurasia, ...

* ''

Vaccinium ''Vaccinium'' is a common and widespread genus of shrubs or dwarf shrubs in the heath family (Ericaceae). The fruits of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry (wh ...

'' * ''

Zanthoxylum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the Family (biology), family Rutaceae that are native to warm temperate and Subtropics, subtropical areas worldwide. It is the type gen ...

References

External links

Engrailed on ''UKMoths''
*
''Lepiforum e.V.''
Boarmiini Moths described in 1775 Moths of Asia Moths of Europe Moths of North America Taxa named by Michael Denis Taxa named by Ignaz Schiffermüller {{Boarmiini-stub

Recorded food plants

References

Further reading

External links