The skuas are a group of predatory
seabird
Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
s with seven species forming the
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Stercorarius'', the only genus in the
family
Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Stercorariidae. The three smaller skuas, the
Arctic skua, the
long-tailed skua, and the
pomarine skua, are called jaegers in
North American English
North American English (NAmE) encompasses the English language as spoken in both the United States and Canada. Because of their related histories and cultures, plus the similarities between the pronunciations (accents), vocabulary, and grammar ...
.
The English word "skua" comes from the
Faroese name for the
great skua, , with the island of
Skúvoy renowned for its colony of that bird. The general Faroese term for skuas is . The word "jaeger" or is German for "hunter".
The genus name ''Stercorarius'' is
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
and means "of dung"; the food disgorged by other birds when pursued by skuas was once thought to be excrement.
Skuas nest on the ground in
temperate
In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
,
Antarctic
The Antarctic (, ; commonly ) is the polar regions of Earth, polar region of Earth that surrounds the South Pole, lying within the Antarctic Circle. It is antipodes, diametrically opposite of the Arctic region around the North Pole.
The Antar ...
, and
Arctic regions, and are long-distance
migrants. They have even been sighted at the
South Pole
The South Pole, also known as the Geographic South Pole or Terrestrial South Pole, is the point in the Southern Hemisphere where the Earth's rotation, Earth's axis of rotation meets its surface. It is called the True South Pole to distinguish ...
.
Biology and habits

Outside the breeding season, skuas take fish,
offal
Offal (), also called variety meats, pluck or organ meats, is the internal organ (anatomy), organs of a butchered animal. Offal may also refer to the by-products of Milling (grinding), milled grains, such as corn or wheat.
Some cultures strong ...
, and
carrion
Carrion (), also known as a carcass, is the decaying flesh of dead animals.
Overview
Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
. Many practice
kleptoparasitism
Kleptoparasitism (originally spelt clepto-parasitism, meaning "parasitism by theft") is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct f ...
, which comprises up to 95% of the feeding methods of wintering skuas, by chasing gulls, terns and other seabirds to steal their catches, regardless of the size of the species attacked (up to three times heavier than the attacking skua). Larger species, such as the
great skua, regularly kill and eat adult seabirds, such as
puffin
Puffins are any of three species of small alcids (auks) in the bird genus ''Fratercula''. These are pelagic seabirds that feed primarily by diving in the water. They breed in large colonies on coastal cliffs or offshore islands, nesting in crev ...
s and
gull
Gulls, or colloquially seagulls, are seabirds of the subfamily Larinae. They are most closely related to terns and skimmers, distantly related to auks, and even more distantly related to waders. Until the 21st century, most gulls were placed ...
s and have been observed killing birds as large as a
grey heron
The grey heron (''Ardea cinerea'') is a long-legged wading bird of the heron family, Ardeidae, native throughout temperate Europe and Asia, and also parts of Africa. It is resident in much of its range, but some populations from the more norther ...
. On the breeding grounds, the three, more slender northern breeding species commonly eat
lemming
A lemming is a small rodent, usually found in or near the Arctic in tundra biomes. Lemmings form the subfamily Arvicolinae (also known as Microtinae) together with voles and muskrats, which form part of the superfamily Muroidea, which also inclu ...
s. Those species that breed in the southern oceans largely feed on fish that can be caught near their colonies. The eggs and chicks of other seabirds, primarily
penguin
Penguins are a group of aquatic flightless birds from the family Spheniscidae () of the order Sphenisciformes (). They live almost exclusively in the Southern Hemisphere. Only one species, the Galápagos penguin, is equatorial, with a sm ...
s, are an important food source for most skua species during the nesting season.
In the southern oceans and
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
region, some skua species (especially the
south polar skua) will readily scavenge carcasses at breeding colonies of both
penguin
Penguins are a group of aquatic flightless birds from the family Spheniscidae () of the order Sphenisciformes (). They live almost exclusively in the Southern Hemisphere. Only one species, the Galápagos penguin, is equatorial, with a sm ...
s and
pinniped
Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant taxon, extant families Odobenidae (whose onl ...
s. Skuas will also kill live penguin chicks and sick or injured adult penguins. In these areas, the skuas will often forfeit their catches to the considerably larger and very aggressive
giant petrels. Skuas have also been observed to directly pilfer
milk
Milk is a white liquid food produced by the mammary glands of lactating mammals. It is the primary source of nutrition for young mammals (including breastfeeding, breastfed human infants) before they are able to digestion, digest solid food. ...
from the
elephant seal's teats.
Skuas are medium to large
bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s, typically with grey or brown plumage, often with white markings on the wings.
[ The skuas range in size from the long-tailed skua, ''Stercorarius longicauda'', at , to the brown skua, ''Stercorarius antarcticus'', at . On average, a skua is about long, and across the wings. They have longish bills with a hooked tip, and webbed feet with sharp claws. They look like large dark gulls, but have a fleshy ]cere
The beak, bill, or Rostrum (anatomy), rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, wikt:grasp#Verb, grasping, and holding (in wikt:probe ...
above the upper mandible.
The skuas are strong, acrobatic fliers. They are generally aggressive in disposition. Potential predators approaching their nests will be quickly attacked by the parent birds, which usually target the heads of intruders – a practice known as ' divebombing'.
Taxonomy
The genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Stercorarius'' was introduced by the French zoologist Mathurin Jacques Brisson
Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher.
Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
in 1760 with the parasitic jaeger (''Stercorarius parasiticus'') as the type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
.
Skuas are related to gull
Gulls, or colloquially seagulls, are seabirds of the subfamily Larinae. They are most closely related to terns and skimmers, distantly related to auks, and even more distantly related to waders. Until the 21st century, most gulls were placed ...
s, wader
245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots
Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
s, auk
Auks or alcids are birds of the family Alcidae in the order Charadriiformes. The alcid family includes the Uria, murres, guillemots, Aethia, auklets, puffins, and Brachyramphus, murrelets. The family contains 25 extant or recently extinct speci ...
s, and skimmer
Skimmer may refer to: Animals
*Skimmer (bird), a common name for birds in the genus ''Rynchops''
*Skimmer (dragonfly), a common name for dragonflies in the family Libellulidae
*Water strider or skimmer, a common name for insects in the family Ge ...
s. In the three smaller species, all nesting exclusively in the Holarctic
The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
, breeding adults have the two central tail feathers obviously elongated, and at least some adults have white on the underparts and pale yellow on the neck. These characteristics are not shared by the larger species, all native to the Southern Hemisphere except for the great skua. Therefore, the skuas are often split into two genera, with only the smaller species retained in ''Stercorarius'', and the large species placed in ''Catharacta''. However, based on genetics
Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinians, Augustinian ...
, behavior, and feather lice, the overall relationship among the species is best expressed by placing all in a single genus. The pomarine and great skuas' mitochondrial DNA
Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
(inherited from the mother) is in fact more closely related to each other than it is to either Arctic or long-tailed skuas, or to the Southern Hemisphere species.[Cohen, Baker, Belchschmidt, Dittmann, Furness, Gerwin, Helbig, de Korte, Marshall, Palma, Peter, Ramli, Siebold, Willcox, Wilson and Zink (1997)]
Enigmatic phylogeny of skuas.
''Proc. Biol. Sci.'' 264(1379):181–190. Thus, hybridization must have played a considerable role in the evolution of the diversity of Northern Hemisphere skuas.
Species
The genus contains seven species:
References
External links
*
Great Skua videos
on the Internet Bird Collection
{{Authority control
Seabirds
Taxa named by Mathurin Jacques Brisson